Last updated: 22 September 2002
Abstract
Abundant new material has been examined mainly from the collections of the National Museum of Namibia (Windhoek) and Natal Museum (Pietermaritzburg). Descriptions of 15 new species and 2 new subspecies, new records for known African species are given. The genus Teuchophorus LOEW is recorded for Tropical Africa for the first time. The Namibian fauna of the family is also briefly reviewed. A key to Namibian genera of the family is presented. Now 25 genera and 41 species of Dolichopodidae are known from Namibia.
Key words: Diptera, Dolichopodidae, Sciapodinae, Medeterinae, new species, new records, keys, Tropical Africa, Namibia.
Introduction
The Afrotropical fauna of Sciapodinae and Medeterinae was recently reviewed by GRICHANOV (1998c, 1999a, 1999b). Treating material from the collections of National Museum of Namibia, Windhoek [NMN], Natal Museum, Pietermaritzburg, South Africa [NMP], the Royal Institute for Natural Sciences, Brussels [RINS] and the Royal Museum for Central Africa, Tervuren [RMCA], I have found abundant new material on the subfamilies Sciapodinae and Medeterinae. Descriptions of 15 new species and 2 new subspecies, new records for known African species are given here. Mesorhaga kirkspriggsi spec. nov., Condylostylus sinclairi spec. nov., Parentia asymmetrica spec. nov., Medetera cimbebasia spec. nov. are described from Namibia, Grootaertia irwini spec. nov., Grootaertia brevipennis spec. nov., Medetera africana africana spec. nov., Medetera calvinia spec. nov., Medetera londti spec. nov., Medetera pallidotiosa spec. nov., Medetera vaalensis spec. nov., Thrypticus parabellus spec. nov. from South Africa, Medetera bweza spec. nov., Corindia demoulini spec. nov. from Congo (Kinshasa), Condylostylus ulrichi spec. nov. from Kenya, Medetera africana senegalensis subspec. nov. from Senegal, Medetera praedator aequatorialis subspec. nov. from Tanzania, Burundi and Congo (Kinshasa). The Genus Teuchophorus LOEW, 1857 is recorded for Tropical Africa for the first time. Now 13 genera and 218 Afrotropical species of Sciapodinae are known. It is the largest subfamily in the Region. Seven genera and 79 Afrotropical species and subspecies of Medeterinae are known from the Region. The subfamily Medeterinae is the most diverse in southern Africa, in contrast to the Sciapodinae with its great number of species in central Africa. The two subfamilies include more than a half of the known dolichopodid species. The Namibian fauna of the family is also briefly reviewed. A first key to Namibian genera of the family is presented. Now 25 genera and 41 species of Dolichopodidae are known from Namibia. Species of the subfamily Medeterinae (13) and genus Medetera FISCHER VON WALDHEIM (10) dominate in the dolichopodid fauna of the country.
Deposition of types of the new species is mentioned under the new names. Bibliography includes works published after the Catalogue of the Diptera of the Afrotropical Region (DYTE & SMITH 1980). Namibian material examined is deposited in the National Museum of Namibia, Windhoek.
DESCRIPTIONS
SUBFAMILY SCIAPODINAE (ZELLER) BECKER, 1917
Genus Mesorhaga SCHINER, 1868
1. Mesorhaga kirkspriggsi GRICHANOV, spec. nov. (Fig. 1)
Holotype. G, Namibia: Rundu dist., Simanya Okawango River, 17°33'17S, 18°32'30E, 23-24.I.1998, KIRK-SPRIGGS & MARAIS, Malaise traps, riverine forest / Mesorhaga spec., det. B.J. SINCLAIR, 1999 [NMN].
Paratype. G with the same 1st label.
Description. Male. Vertex deeply excavated. Frons and face metallic dark-green, whitish pollinose. A fine front vertical seta bends forward on each side; ocellar tubercle with 1 pair of strong bristles and pair of weaker posterior setae; 2 long postvertical setae positioned as a linear continuation of the postocular setal row. Upper postocular setae black, lateral postoculars white. Ventral postcranium covered with irregular white hairs. Face narrowed, approximately 1.3 times as high as wide under antennae. Proboscis and palpus brown, with short light hairs; palpus with 1 strong black seta in addition. Antennae black. Pedicel with ring of short setae; 1 or 2 ventral setae longer than pedicel. First flagellomere rounded, as long as high, with short hairs. Arista microscopically haired, with 1st segment slightly thickened. Length ratio of scape to pedicel to first flagellomere to segments of arista, 5 : 4 : 6 : 5 : 37.
Mesonotum and scutellum metallic bluish-green. Pleura bronze-black, whitish pollinose. 5 strong dorsocentral setae, 1-2 pairs of weak acrostichals anteriorly, a pair of strong and pair of fine lateral scutellars, half as long as medial setae.
Legs mostly yellow; fore coxa yellow, middle coxa mostly dark-brown, hind coxa brown in basal half, hind tibia with narrow brown stripe on posterodorsal surface in the 2nd quarter, apical segments of fore tarsus brown, mid tarsus black from tip of basitarsus, hind tarsus black from 2nd half of basitarsus. Fore and middle coxae with numerous yellow cilia anteriorly; hind coxa with yellow external seta and several hairs. All femora with light ventral hairs, shorter than diameter of femora. Fore tibia bare; last tarsomere of fore tarsus slightly flattened. Length ratio of fore coxa to femur to tibia to tarsus (segments from first to fifth), 30 : 57 : 57 : 30 : 13 : 9 : 6 : 7. Middle tibia and basitarsus with several apical setae, tarsus simple. Length ratio of middle coxa to femur to tibia to tarsus (segments from first to fifth), 25 : 65 : 74 : 44 : 18 : 12 : 10 : 7. Hind tibia without strong setae. Last tarsomere of hind tarsus slightly flattened. Length ratio of hind coxa to femur to tibia to tarsus (segments from first to fifth), 20 : 70 : 85 : 30 : 28 : 18 : 12 : 8.
Wings yellowish; veins pale brown. R1 reaching mid-wing. R2+3 straight. R4+5 gently curved to M1 at apex. M1 with two 135° bends; the basal bend positioned at 1st third of distal part of the vein. Ratio of parts of costa between R2+3 and R4+5 to those between R4+5 and M1, 33 : 7. M2 absent. Crossvein m-cu straight. Ratio of crossvein m-cu to apical part of M1+2 (up to curvation) to apical part of CuA1, 18 : 28 : 40. Alula weakly developed. Fold-like anal vein and broad anal lobe present. Anal angle acute or right. Lower calypter yellow with brownish rim and fan of dark cilia. Halter pale yellow.
Abdomen bronze-black, with short black hairs. First tergite with several long black distolateral bristles. Ventrum with white hairs. Seventh segment shortened. Hypopygium entirely black-brown, epandrium elongate. Cercus hook-like, strongly curved ventrad, thick in basal half, with short dorsolateral setae, narrowed beyond middle, widened at apex; distal part with distinct dorsal process and several simple setae as figured. Surstylus bi- or trilobate, with several distal setae. Epandrial lobe long, flattened, bearing one strong seta beyond the middle and very small seta at apex.
Female unknown.
Length (mm): body 2.4-2.6; antenna 0.7; wing 2.5/1.0; hypopygium 0.8.
Distribution: Namibia.
Etymology. The species is named for one of the collectors, A.H. KIRK-SPRIGGS.
Diagnosis. The new species is related to M. tsurikovi GRICHANOV 1998c: Fig. 3, differing in smaller size, entirely yellow femora, CuA1 twice longer than crossvein m-cu and hypopygium morphology.
Genus Condylostylus BIGOT, 1859
2. Condylostylus sinclairi GRICHANOV, spec. nov. (Fig. 2)
Holotype. G, Namibia: Rundu dist., Katara Okavango R., 17°48'56S, 18°53'38E, 20-23.I.1998, KIRK-SPRIGGS & MARAIS, Malaise traps / Namibian National Insect Collection, National Museum, P.O. Box 1203, Windhoek, Namibia.
Paratypes [mostly on pins, two of them with additional label: Condylostylus spec., det. B.J. SINCLAIR, 1999]. 7GG, 3EE, same labels; 20GG, 3EE, Namibia: Rundu dist., Simanya Okawango River, 17°33'17S, 18°32'30E, 23-24.I.1998, KIRK-SPRIGGS & MARAIS, Malaise traps, riverine forest; 4GG, 1E, Namibia: Rundu dist., 1 km S of Katara, 17°50'25S, 18°54'26E, 22-23.I.1998, KIRK-SPRIGGS & MARAIS, Malaise traps, primary forest; 1G, 1E, Namibia: Rundu dist., Matende River, 17°54'46S, 19°35'16E, 20-21.I.1998, MARAIS & KIRK-SPRIGGS, Malaise traps, primary forest; 1G [in alcohol], Leeupan, Kaudom Game reserve, 18°40'S, 20°52'E, 12-14.I.1991, E. MARAIS; 2GG, 1E [in alcohol], Mangetti, Quarantine Camp: Kavango, 18°40'S, 19°02'E, 11.I.1993, E. MARAIS [NMN].
Description. Male. Frons metallic blue-green, shining. A strong front vertical bristle bends forward, arising from small mound covered with 8-10 white hairs; fine postvertical bristle is positioned as a linear continuation of the postocular setal row. Ocellar tubercle with a pair of strong setae and pair of posterior hairs. Upper postocular setae short, black, uniseriate; lower postoculars white, in several rows. Ventral postcranium covered with irregular white hairs. Face greenish-blue, silvery-white pollinose, broad; ratio of its height to width under antennae to width at clypeus, 55 : 36 : 20. Bulging clypeus nearly as high as epistome. Proboscis brownish, palpi black, with light hairs and 1-3 black setae. Antennae black, slightly longer than height of head, with small simple segments. Pedicel with short setulae laterally, 1-2 dorsal and 2-3 ventral very long bristles, which 2-3 times longer than 1st flagellomere. First flagellomere rounded, not longer than high, with short hairs. Arista dorsal, microscopically haired. Length ratio of scape to pedicel to first flagellomere to arista, 6 : 6 : 6 : 95.
Mesonotum and scutellum metallic bluish-green, slightly pollinose. Pleura bronze, white pollinose. Five dorsocentral bristles gradually decreasing in size anteriorly with two strongest posterior pairs. Usually 2 pairs of long acrostichals. Scutellum with two pairs of strong setae.
Legs mostly yellow. Mid coxa with brown external spot; hind femur brownish at extreme apex; tarsi black except for base of anterior four basitarsi. Fore coxa from the front with numerous yellow hairs and 3 black subapical setae. Mid coxa from the outside with light hairs and several black cilia. Hind coxa with one black external seta and several yellow hairs. Femora with 2 rows of white fine ventral hairs, slightly longer than femora diameter. Fore tibia with 2 long black apical posteroventral setae. Fore basitarsus slightly swollen in basal half, with ventral pile along entire length. Length ratio of fore coxa to femur to tibia to tarsus (segments from first to fifth), 50 : 80 : 92 : 78 : 31 : 19 : 12 : 9. Middle tibia with 1 anterodorsal at basal fifth and several apical setae, without remarkable hairs. Mid tarsus simple. Length ratio of middle coxa to femur to tibia to tarsus (segments from first to fifth), 40 : 96 : 135 : 110 : 28 : 21 : 10 : 10. Last tarsomeres of hind tarsi slightly thickened. Length ratio of hind coxa to femur to tibia to tarsus (segments from first to fifth), 30 : 114 : 177 : 84 : 30 : 18 : 12 : 8.
Wing elongate-oval, mostly hyaline, brown along costa in apical 2/3, with 2 broad transverse brown stripes along m-cu and vein-fork M1+2; veins brown. Venation undisturbed. R4+5 gently curved to M1 in apical fifth. M1+2 slightly convex anteriad in apical half. M1 strongly convex basad, forming acute angle with M1+2. M2 as a linear continuation of M1+2. Ratio of part of costa between R2+3 and R4+5 to this between R4+5 and M1, 49 : 5. Crossvein m-cu straight. Ratio of crossvein m-cu to apical part of M1+2 (fork-handle) to apical part of CuA1, 38 : 65 : 29. Anal vein fold-like, anal lobe and alula developed. Anal angle acute. Lower calypter brown, with mostly black cilia. Halter brown; halter stem thin, 1.5 times longer than knob, with row of setulae in front of knob.
Abdomen thin and long, metallic green-blue-black, posteriorly entirely black-violet, with short black hairs and long fine marginal setae. First tergite with membranous excavation, longitudinal dorsal furrow and white lateral hairs; ventrum with white hairs. Unmodified segments combined 2 times as long as mesonotum. 5-6th segments swollen, 7th tergite short. Hypopygium subtriangular, black, with short black hairs. Hypandrium distinct. Cercus long, filiform, slightly swollen in basal 1/3, black, with numerous short black hairs along entire length. Cercus 2 times as long as epandrium. Surstylus and epandrial lobe greatly reduced.
Female similar to male except lacking male secondary sexual characters.
Length (mm): body 4.4; antenna 1.4; wing 4.7/1.6; postabdomen 0.9.
Distribution: Namibia.
Etymology. The species is named for Canadian dipterologist, Dr. B.J. SINCLAIR.
Diagnosis. The new species is close to C. paricoxa PARENT 1939 as redescribed by GRICHANOV (1996a: Fig. 3), differing in darker tarsi, wider face, longer fore basitarsus, black antenna, long black cercus etc. Frons with white hairs; fore tibia with 2 long black apical posteroventral setae. First tarsomere slightly swollen in basal half, with short dense ventral hairs. Afrotropical Condylostylus species can be distinguished mainly by weak variations of male cercus shape in combination with leg setation and coloration. Surstyli are greatly reduced in African species (in contrast to American and Oriental species) and have no diagnostic value.
3. Condylostylus ulrichi GRICHANOV, spec. nov. (Fig. 3)
Holotype [in alcohol]. G, Kenya, Taita Hills, Macha, 8.I-14.I.1999 [RMCA].
Description. Male. Frons metallic blue-green, shining. A strong front vertical bristle bends forward, arising from small bare mound; strong postvertical bristle is positioned as a linear continuation of the postocular setal row. Ocellar tubercle with a pair of strong setae. Ventral postcranium covered with irregular white hairs. Face greenish-blue, mat, narrow; 10 times as high as wide in the middle and 3.3 times as high as wide under antennae. Bulging clypeus 2/3 as wide as epistome under antennae. Proboscis brown, palpi black, with light hairs; palpus also with 2 black cilia. Antennae black, nearly as long as height of head. Pedicel with several dorsal and ventral bristles, which not much longer than 1st flagellomere. First flagellomere subtriangular with rounded apex, as long as or slightly longer than high, with short hairs. Arista positioned just before middle of dorsal side, microscopically haired. Length ratio of scape to pedicel to first flagellomere to arista, 5 : 5 : 8 : 65.
Mesonotum and scutellum metallic blue-green. Pleura bronze-black. Five dorsocentral bristles gradually decreasing in size anteriorly. Short acrostichals in two rows, restricted to anterior 2/3 of mesonotum. Scutellum with a pair of strong medial setae and two lateral setae, 2/3 as long as medial.
Legs mostly yellow. Fore coxa yellow, mid and hind coxae almost entirely black; hind femur with brownish anterodorsal stripe, almost entirely brown at apex; hind tibia blackish-brown; fore tarsus brown from middle of basitarsus, mid tarsus mostly black except for brownish basitarsus in its basal half, hind tarsus black. Fore coxa from the front with numerous yellow hairs and 1 black subapical seta. Mid coxa from the outside with light hairs. Hind coxa with one fine light external seta and several yellow hairs. Femora with white fine ventral hairs at base, at most half as long as femora diameter. Fore tibia with one long semierect subapical ventral seta. Fore basitarsus 10 times longer than wide, slightly flattened and widened in distal half, with ventral pile and row of elongate posteroventral setulae in basal 1/2. Length ratio of fore coxa to femur to tibia to tarsus (segments from first to fifth), 60 : 85 : 108 : 78 : 25 : 15 : 10 : 8. Middle tibia inconspicuously swollen; ventrally free of setulae, with pile of microscopic hairs; with posteroventral row of elongate setulae along entire length, longer than tibia diameter, and 3 apical setae. Fourth and fifth tarsomeres slightly enlarged and flattened, with elongate setulae. Length ratio of middle coxa to femur to tibia to tarsus (segments from first to fifth), 40 : 108 : 165 : 112 : 25 : 18 : 11 : 20. Hind tarsus simple. Length ratio of hind coxa to femur to tibia to tarsus (segments from first to fifth), 30 : 145 : 215 : 95 : 32 : 21 : 15 : 11.
Wing long and narrow, mostly hyaline, brownish anteriorly in distal 1/3; veins brown. Venation undisturbed. R4+5 gently curved to M1 in apical fifth. M1+2 straight. M1 with elbow-like curvation, forming right angle with M1+2. M2 distinct. Ratio of part of costa between R2+3 and R4+5 to this between R4+5 and M1, 35 : 4. Crossvein m-cu straight. Ratio of crossvein m-cu to apical part of M1+2 (fork-handle) to apical part of CuA1, 34 : 65 : 17. Anal vein and lobe reduced. Anal angle absent. Lower calypter brown, with light cilia. Halter yellow; halter stem thin, 1.5 times longer than knob, with row of setulae in front of knob.
Abdomen thin and long, metallic green-blue-black, posteriorly entirely black-violet, with short black hairs and setae. First tergite with membranous excavation, longitudinal dorsal furrow and white lateral hairs. Unmodified segments combined 3 times as long as mesonotum. 5-6th segments swollen, 7th tergite short. Hypopygium black, with short black hairs. Cercus long, strip-like, black, narrowed and brownish in distal 1/3, with numerous short, ventrally hooked hairs along entire length. Cercus nearly 3 times as long as epandrium. Surstylus and epandrial lobe greatly reduced.
Female unknown.
Length (mm): body 5.2; antenna 1.0; wing 4.3/1.2; hypopygium 1.3.
Distribution: Kenya.
Etymology. The species is named for German dipterologist, Dr. H. ULRICH.
Diagnosis. The new species is close to C. basovi GRICHANOV 1998c: Fig. 8, differing in mostly yellow legs, narrow fore basitarsus, ornamented mid tibia, shorter cercus and other characters. See also diagnosis of C. sinclairi.
Genus Parentia HARDY, 1935
4. Parentia asymmetrica GRICHANOV, spec. nov. (Figs. 4-5)
Holotype [in alcohol]. G, Namibia: West Caprivi Park, Kwando River: Susuwe, 17°45'37S, 23°20'55E, 28.IX.-2.X.1998, A.H. KIRK-SPRIGGS, Malaise trap, dry woodland [NMN].
Paratypes [in alcohol]. 5GG, 6EE, same label.
Description. Male. Frons metallic blue-green, shining. A strong front vertical bristle bends forward, arising from small bare mound; shorter postvertical bristle is positioned as a linear continuation of the postocular setal row. Ocellar tubercle with a pair of strong setae and pair of hairs. Ventral postcranium covered with irregular white hairs. Face (including clypeus) bluish-green, narrowed towards palpi, 2 times as high as wide at clypeus and 1.2 times as high as wide under antennae. Weakly bulging clypeus not reaching lower margin of eyes. Proboscis brown, palpus brown, with light hairs and 2 black setae. Antennae black, slightly longer than height of head. Pedicel globular, with 1-2 long dorsal and ventral bristles, 2 times longer than pedicel. First flagellomere rounded, as long as high, with short hairs. Arista dorsal, microscopically haired. Length ratio of scape to pedicel to first flagellomere to arista (1st to 2nd segments), 5 : 5 : 5 : 4 : 86.
Thorax metallic blue-green. Five or six dorsocentral bristles gradually decreasing in size anteriorly and 1-2 anterior hairs. Usually three pairs of long acrostichals, nearly as long as anterior dorsocentrals. Scutellum with two pairs of strong setae with lateral setae 2/3 the length of medial.
Legs mostly yellow. Fore coxa yellow, middle coxa dark-brown, hind coxa brown, hind femora with brownish dorsal spot at extreme apex; hind tibia brown at extreme apex, anteriorly at basal 1/5 with small brown areole having microscopic setulae situated somewhat thicker than usually; fore and mid tarsi black from tip of basitarsus, hind tarsus entirely black. Fore coxa from the front with numerous yellow hairs and 3 yellow subapical setae. Middle coxa from the outside with white hairs and cilia. Hind coxa with several white hairs in addition to one yellow external seta. Fore femur with short yellow hairs ventrally. Fore tibia with 3-4 short apical seta. Fore tarsus simple. Length ratio of fore coxa to femur to tibia to tarsus (segments from first to fifth), 35 : 69 : 73 : 49 : 19 : 11 : 8 : 6. Mid femur with 1 yellow ventral seta at base, as long as femur diameter, ventral row of black cilia in distal half, half as long as femur diameter. Mid tibia with 4-5 apical setae. Mid tarsus practically simple, basitarsus with elongate setulae slightly longer than diameter of tarsomere and areole in distal 1/2 of ventral side free of setulae. Length ratio of middle coxa to femur to tibia to tarsus (segments from first to fifth), 28 : 85 : 108 : 60 : 23 : 15 : 8 : 7. Hind tibia with a callus described above, several inconspicuous dorsal and 2-3 short apical setae. Hind tarsus simple. Length ratio of hind coxa to femur to tibia to tarsus (segments from first to fifth), 20 : 100 : 123 : 52 : 28 : 17 : 10 : 8.
Wings elongate-oval, mostly greyish with posterior margin widely hyaline; veins brown. Venation undisturbed. R1 reaching 2/5 of wing length. R4+5 gently curved to M1 in apical fifth. M1+2 straight. M1 with wide arc, forming right angle with M1+2. M2 as faint fold on membrane. Ratio of part of costa between R2+3 and R4+5 to this between R4+5 and M1, 19 : 5. Crossvein m-cu straight. Ratio of crossvein m-cu to apical part of M1+2 (fork-handle) to apical part of CuA1, 32 : 33 : 22. Anal vein foldlike, anal lobe and alula developed. Anal angle acute. Lower calypter yellow-brownish, with fine light cilia. Halter yellow, halter stem thin, 1.5 times longer than knob, with row of setulae in front of knob.
Abdomen metallic green, with short black hairs and marginal setae. First tergite with narrow membranous excavation, longitudinal dorsal furrow, two pairs of long black dorsolateral setae and short white lateral hairs. Unmodified segments combined 2 times as long as thorax. Hypopygium brown-black, with black hairs. Hypandrium basoventral, long, hood-like. Aedeagus with double dorsal angle. Cercus yellow-brownish, short, slightly swollen at base, narrow at apex, with numerous black hairs along entire length and several long undulate apical setae. Surstyli and epandrial lobes asymmetric. Left surstylus short and thin, spine-shaped; right surstylus long and broad, nearly as long as cercus. Left epandrial lobe as thin curved process with several short subapical setae; right epandrial lobe as broad projection with 2 apical setae.
Female. Similar to male except lacking male secondary sexual characters. Hind coxa, femur and tibia entirely yellow, hind basitarsus yellow at base. Femora bare; fore tibia with 1 anterodorsal, 1 posterodorsal short setae; mid tibia with 2 anterodorsal, 2 posterodorsal setae; hind tibia with 2 anterodorsal, 4 posterodorsal setae; hind basitarsus with small basoventral seta; 1st abdominal tergum without long setae.
Length (mm): body 3.6-4.4; antenna 0.9; wing 3.5-4.0/1.0; hypopygium 0.85.
Distribution: Namibia.
Diagnosis. P. asymmetrica is closely related to P. stenurus (LOEW 1858) (described also by CURRAN as Condylostylus sicatrix CURRAN 1926), differing in almost entirely yellow hind tibia and practically simple middle leg. The new species differs from all other known species of the genus in remarkably asymmetric hypopygium. Male wing has no diagnostic features, being similar to female wing. P. asymmetrica is the most close to Parentia and cannot be associated with Condylostylus, although many Australian species of the genus has quite different male secondary sexual characters (MSSC). Afrotropical species of the genus are confined to southern Africa.
SUBFAMILY MEDETERINAE (FISCHER VON WALDHEIM) LIOY, 1864
Genus Corindia Bickel, 1986
5. Corindia demoulini GRICHANOV, spec. nov. (Fig. 6)
Holotype. Male, Congo Belge, P.N.G., Miss. DE SAEGER, II/gd/16, 26-IX-1952, Rec. H. DE SAEGER, 4084 [RMCA].
Paratypes. 14 males with the same label as holotype (one of them collected by G. DEMOULIN and one by J. VERSCHUREN), differing in the collection dates and codes as follows: I/c/2, 23-VIII-1950, 768; II/gd/10, 28-XII-1951, 2954; II/gd/4, 18-I-1952, 3024; II/gd/6, 22.I.1952, 3031; II/gd/11, 11-III-1952, 3183; II/gd/11, 24-VI-1952, 3701; II/gd/6, 2.IX.1952, 4023; II/gd/16, 26-IX-1952, 4084 [RMCA].
Description. Male. Frons blue-violet. Face entirely shining blue-green. One strong vertical seta laterally of ocellar tubercle; the latter is low and broad, with 2 strong setae and 2 hairs; one fine postvertical seta, one row of short postocular setae present. Face widest under antennae, narrowed downward, clypeus parallel-sided. Ratio of height of epistome to its maximal width to height of clypeus , 18 : 14 : 9. Antenna 1/4 longer than head; scape and pedicel reddish-yellow, 1st flagellomere black-brown; scape vase-like; pedicel globular, with a ring of short apical setulae; first flagellomere smaller than pedicel, as long as high, irregularly rounded. Arista apical, with very short basal segment and short hairs, 6 times longer than antennomeres combined. Palpus and proboscis short, black-brown, with black hairs; palpus with one black seta.
Mesonotum flattened in posterior third, metallic blue-green, with light-brown setae. Pleura metallic green. Five pairs of strong dorsocentral setae decreasing in size anteriorly. Acrostichal setae biseriate, becoming quadriseriate on mesonotal flattening. One light propleural seta. Scutellum with a pair of strong and a pair of short hairlike lateral setae.
Legs mostly yellow; fore coxa yellow-brownish at base, middle and hind coxae brown with yellow apex, last tarsomeres of all tarsi brown. Fore coxa with short hairs and several apical setae; middle coxa with hairs and one external seta; hind coxa with one strong and one fine external setae. Fore leg without setae. Middle tibia with one strong anterodorsal at basal fourth and one strong apicoventral setae. Hind femur simple. Hind tibia with short setae. All tarsi simple. Length ratio of fore coxa to femur to tibia to tarsus (segments from first to fifth), 35 : 47 : 43 : 24 : 8 : 5 : 4 : 6. Same ratio for middle leg, 30 : 55 : 53 : 28 : 18 : 12 : 6 : 5. Same ratio for hind leg, 20 : 60 : 70 : 15 : 26 : 20 : 12 : 8.
Wing simple, hyaline, veins brown; posterior wing margin evenly convex; maximum wing-width at the end of CuA1. Costa without long hairs, with microsetulae reaching middistance between ends of R2+3 and R4+5. R1 reaching to first fifth of wing. R2+3 almost straight. R4+5 inconspicuously convex anteriad. Ratio of part of costa between R2+3 and R4+5 to this between R4+5 and M1+2, 36 : 8. Ratio of apical to basal part of M1+2, 82 : 51. R4+5 and M1+2 weakly convergent, almost parallel in apical part. Ratio of cross-vein m-cu to maximal distance between R4+5 and M1+2 to apical part of CuA1, 17 : 12 : 30. Anal vein absent; anal lobe small. Alula undeveloped. Lower calypter yellow, with brownish cilia. Halter yellow.
Abdomen metallic dark-bluish-green, with short light setae. Hypandrium, surstylus and cercus yellow. Epandrium black-green, elongate-triangular, with basolateral foramen. One epandrial seta at base of hypandrium. Epandrial lobe long, broad, with 2 long setae at apex. Hypandrium long and thin, swollen at apex. Aedeagus thin, broadened at apex. Surstylus simple, with about 6 strong apical setae. Cercus suboval; cercus and thin distolateral cercal arm with long dorsal setae as figured.
Female unknown.
Length (mm): body without antennae 2.0-2.3, antenna 0.9, wing 2.2/0.8, postabdomen 0.8.
Distribution: Congo (Kinshasa).
Etymology. The species is named for one of the collectors, G. Demoulin.
Diagnosis. The new species is close to C. danielssoni GRICHANOV 1998d: Fig. 3, differing in larger size, several strong apical setae on surstylus and other characters of hypopygium. Other Afrotropical species of Corindia have been collected also in the Garamba National Park that has mainly savanna landscape with humid gallery forests along rivers only. The Australian species of the genus are characteristic of sclerophil Eucalyptus forests. To date none species of Corindia was recorded in rainforests.
Genus Grootaertia GRICHANOV, 1999
6. Grootaertia irwini GRICHANOV, spec. nov. (Fig. 7)
Holotype. G, South Africa, Cape Prov., 14 mi. SE Langebaan, 200 ft, Sept. 18, 1972, M.E. IRWIN, 3318Aa, coastal dunes and sandy plain [NMP].
Paratypes. 13 GG and 12 EE with the same label as holotype.
Description. Male. Body generally black, with blackish, though shining light setae. Frons and face black, grey pollinose. Ocellar tubercle prominent, with a pair of strong setae. One strong vertical seta laterally on frons present, a strong postvertical one is positioned as a linear continuation of the postocular setal row; postocular setae blackish above, white laterally and below. Ventral postcranium with several long cilia. Face widest under antennae, gradually narrowed towards palpi. Clypeal suture distinct. Ratio of height of face to its maximal width, 20 : 10. Antenna as long as head height, black; pedicel with a ring of short apical setulae; 1st flagellomere elongate-triangular, 1.5 times longer than high, with acute apex and very short hairs. Arista apical, almost glabrous, as long as antennomeres combined. Length ratio of scape to pedicel to first flagellomere to arista, 3 : 4 : 14 : 21. Palpus and proboscis black, with sparse hairs; palpus short, with 1 black seta; proboscis stout, prominent.
Thorax entirely black, grey pollinose, mesonotum concave in posterior third. Four pairs of strong dorsocentral setae with fourth seta shortened and additional small seta in front of the 1st one. Acrostichal setae absent. One long and one very short humeral, one posthumeral, 2 strong notopleural, 1-2 short presutural and 2 short and strong postsutural (supraalar) setae present. Two propleural setae. Scutellum with a pair of strong setae.
Legs with all coxae and femora blackish-brown, tibia and basitarsi dirty-yellow or brownish, other tarsomeres black. Fore coxa with short hairs and several light setae in apical half; mid and hind coxae with one external seta. Femora without setae and long hairs. Fore tibia with 2-3 short apicoventral setae. Mid tibia with 1 anterior and 1 posterodorsal setae at basal 1/5, 2-3 short apical setae in addition to 1 strong apicoventral seta. Hind tibia with several short dorsal setae; hind basitarsus with 1 short basoventral seta. Length ratio of fore coxa to femur to tibia to tarsus (segments from first to fifth), 21 : 29 : 33 : 16 : 7 : 5 : 4 : 5. Same ratio for middle leg, 15 : 38 : 45 : 22 : 9 : 6 : 5 : 6. Same ratio for hind leg, 12 : 36 : 45 : 15 : 12 : 8 : 6 : 5.
Wing elongate-oval, simple, hyaline; veins brown; posterior wing margin evenly convex; maximum wing-width just before the end of CuA1. Costa without long hairs. Ratio of part of costa between R2+3 and R4+5 to this between R4+5 and M1+2, 16 : 6. R1 reaching 2/5 of wing length. R2+3, R4+5 and M1+2 slightly convex anteriorly. R4+5 and M1+2 parallel in apical part. Ratio of apical to basal part of M1+2 (from r-m), 55 : 25. Crossvein m-cu slightly convex. Ratio of cross-vein m-cu to maximal distance between R4+5 and M1+2 to apical part of CuA1, 7 : 7 : 28. Anal vein fold-like; anal lobe poorly developed; anal angle obtuse. Alula reduced. Lower calypter small, brown, with greatly reduced cilia. Halter black.
Abdomen black, weakly pollinose, with short setae. 7th tergum semicircular, narrow, lying conformably with 6th tergum. 8th tergum large, covering left basolateral foramen. Epandrium shining black-brown, elongate-oval (lateral view). No epandrial seta. Hypandrium (epandrial lobe?) as long pointed lobe positioned at left distoventral angle of epandrium. Aedeagus long, widened in middle; lateral lobes of aedeagus long, thin, pointed apically. Surstyli brown, glabrous, two pairs, positioned apicodorsally. Cercus brown, visible, with several long dorsal setae.
Female similar to male except lacking male secondary sexual characters. 1st flagellomere slightly longer than high; arista 1.6 times longer than antennomeres combined.
Length (mm): body without antennae 1.4, antenna 0.5, wing 1.5/0.6, hypopygium 0.6.
Distribution: South Africa.
Etymology. The species is named after the American dipterologist, Dr. M.I. IRWIN, who has collected a type series of the new species.
Diagnosis. The new species differs from other species of the genus in entirely black body, blackish coxae and femora. It is close to G. kuznetsovi GRICHANOV 1999a: Fig. 15, strongly differing in morphology of hypopygium. G. kuznetsovi has long dorsal setae on ventral surstyli, large cercus with distinct fingerlike distolateral lobe.
7. Grootaertia brevipennis GRICHANOV, spec. nov. (Fig. 8)
Holotype. G, S. Africa: Cape, #51, Hermanus-Heys Hill, 34°25'S, 19°14'E, 60 m, Date: 7.X.1993, Coll. J.G.H. LONDT, Flowers & Dassie hole [NMP].
Description. Male. Frons black, grey pollinose; face light-brown, white pollinose. Ocellar tubercle prominent, with a pair of strong black setae and 2 short hairs. One strong brown vertical seta laterally on frons present, a strong postvertical one is positioned as a linear continuation of the postocular setal row; postocular setae white. Ventral postcranium with several long cilia. Face widest under antennae, slightly narrowed towards palpi. Clypeal suture marked laterally. Antenna shorter than head height; scape and pedicel yellow; pedicel with a ring of short apical setulae; 1st flagellomere black, rounded, flattened laterally, as long as high, with very short hairs. Arista apical, glabrous, twice longer than antennomeres combined. Length ratio of scape to pedicel to first flagellomere to arista, 4 : 4 : 7 : 34. Palpus and proboscis yellow-brownish, with sparse hairs; palpus short, with 1 light seta; proboscis stout, prominent.
Mesonotum concave in posterior third, mostly yellow-brownish, grey pollinose, with dark-brown setae. Mesonotal depression and scutellum bluish-black dorsally; mesonotum anteriorly with medial and two lateral (along rows of dorsocentral setae) dark narrow longitudinal stripes. Pleurae reddish-yellow, with brown longitudinal medial stripe. Four pairs of strong dorsocentral setae with 1 additional small seta in front of the 1st one. Acrostichal setae absent. One long and one very short humeral, one posthumeral, 1 long and strong notopleural, 2 short presutural and 2 strong postsutural (supraalar) setae present. Propleura with one light seta and 2-3 cilia above. Scutellum with a pair of strong setae.
Legs including coxae yellow; 5th segment of all tarsi black. Fore coxa with short hairs and several apical setae; mid and hind coxae with one external seta. Femora without setae and long hairs. Fore tibia with 2-3 short apical setae. Mid tibia with 1 anterior and 1 posterodorsal setae at basal 1/4, 2-3 short apical setae in addition to 1 strong apicoventral seta. Hind tibia with 2 short dorsal setae, hind basitarsus with 1 short basoventral seta. Length ratio of fore coxa to femur to tibia to tarsus (segments from first to fifth), 25 : 42 : 45 : 24 : 10 : 7 : 5 : 6. Same ratio for middle leg, 20 : 45 : 53 : 30 : 11 : 8 : 5 : 6. Same ratio for hind leg, 15 : 50 : 62 : 21 : 15 : 10 : 6 : 7.
Wing elongate-oval, simple, hyaline; veins yellow-brown; posterior wing margin evenly convex; maximum wing-width just before the end of CuA1. Costa without long hairs. Ratio of part of costa between R2+3 and R4+5 to this between R4+5 and M1+2, 19 : 9. R1 reaching 2/5 of wing length. R2+3, R4+5 and M1+2 almost straight, slightly convex anteriorly. R4+5 and M1+2 parallel in apical part. Ratio of apical to basal part of M1+2 (from r-m), 85 : 32. Crossvein m-cu slightly convex. Apical part of CuA1 slightly concave, fold-like at wing margin. Ratio of cross-vein m-cu to maximal distance between R4+5 and M1+2 to apical part of CuA1, 11 : 8 : 42. Anal vein fold-like; anal lobe developed; anal angle obtuse. Alula undeveloped. Lower calypter yellow, with greatly reduced cilia. Halter yellow.
Abdomen mostly dark-brown, weakly pollinose, with short setae. Sterna yellow, weakly sclerotized. 7th tergum semicircular, narrow, lying conformably with 6th tergum. 8th tergum large, covering left basolateral foramen. Epandrium elongate (lateral view). No epandrial seta. No epandrial lobe. Hypandrium large, almost rectangularly curved, positioned on left side of epandrium. Aedeagus short and broad, with thin apical part; lateral lobes of aedeagus asymmetric. Surstyli yellow, short, hook-like, glabrous; one pair of surstyli attached to epandrium dorsoapically. Cercus large, exposed, with several dorsal setae.
Female unknown.
Length (mm): body without antennae 1.8, antenna 0.6, wing 2.0/0.7, hypopygium 0.6.
Distribution: South Africa.
Diagnosis. The new species is similar to G. asymmetrica GRICHANOV 1999a: Fig. 9, reliably differing from the latter species in hypopygium morphology only. G. asymmetrica has hypandrium short, spade-like, slightly widened distally, with widely rounded distal margin (ventral view), apicoventral in position; one pair of long narrow glabrous pointed surstyli, approximately as long as epandrium: left surstylus simple, dorsolateral in position, arising at base of cercus; right surstylus ventrolateral in position, arising at base of hypandrium, with narrow basal process 1/3 as long as surstylus.
Genus Medetera FISCHER VON WALDHEIM, 1819
8. Medetera africana africana GRICHANOV, spec. nov. (Fig. 9)
Holotype. G, Scottburg, Natal, S. Africa, B. & P. STUCKENBERG, 15.XI.1963 [NMP].
Paratype. G, South Africa, Natal, Zululand, Ndumu-Game Reserve, 26.X.1972, ME IRWIN, 2632Cc [NMP].
Description. Male. Frons and face black, grey pollinose; clypeus weakly shining in middle. A row of several short black postocular setae at the top of eye and a row of light-brownish postoculars below present; two lowest postoculars black. One strong vertical seta laterally on frons. Ocellar tubercle with one pair of strong setae. Ventral postcranium covered with sparse light irregular setae. Face broad, widest under antennae, narrowest at middle, slightly widened below middle. Ratio of height of epistome to its maximal width to height of clypeus to its minimal width, 23 : 17 : 14 : 14. Antenna black, nearly as long as head height; pedicel slightly swollen, with a ring of short apical setulae; first flagellomere small, rounded, with distinct apex and short terminal hairs. Arista subapical, microscopically pubescent. Length ratio of scape to pedicel to first flagellomere to arista, 4 : 5 : 6 : 55. Palpus and proboscis short, black, weakly pollinose, with light hairs; palpus with one black seta.
Thorax bronze-black, with bluish reflection, grey pollinose, with black setae. Two pairs of strong and two pairs of very short dorsocentral setae with several microscopic hairs anteriorly. Two rows of short acrostichals extending to mesonotal flattening. Propleura with 1 strong black seta and 1 fine hair above fore coxa. Scutellum with a pair of strong medial setae and two lateral setae, half as long as medial.
Legs mostly dirty-yellow, all coxae brown, femora slightly darkened in basal half, apical segments of tarsi brown. Coxae with brownish setae and hairs; fore coxa with short hairs and several apical setae; mid and hind coxae each with one external seta. Fore legs without setae. Fore tarsus simple. Length ratio of fore coxa to femur to tibia to tarsus (segments from first to fifth), 43 : 65 : 65 : 27 : 25 : 17 : 10 : 9. Mid tibia with one anterodorsal, one posterodorsal at basal 1/4 and 3-4 apical setae. Length ratio of mid coxa to femur to tibia to tarsus (segments from first to fifth), 35 : 70 : 79 : 38 : 32 : 20 : 9 : 8. Hind legs without long setae. Hind tibia posteriorly with several short black spinules covered with apical scale. Hind tarsus simple. Length ratio of hind coxa to femur to tibia to tarsus (segments from first to fifth), 27 : 75 : 94 : 17 : 48 : 22 : 11 : 9.
Wings hyaline; veins brown, yellow at base. Costa without long hairs. Ratio of part of costa between R2+3 and R4+5 to this between R4+5 and M1+2, 19 : 6. R4+5 and M1+2 distinctly convergent in apical part. Ratio of apical to basal part of M1+2 (from r-m), 10 : 7. Ratio of cross-vein m-cu to maximal distance between R4+5 and M1+2 to apical part of CuA1, 16 : 22 : 17. Lower calypter yellow, with light cilia. Halters yellow.
Abdomen black-brown, pollinose, with short black setae; 7th segment half as long as epandrium. Epandrium elongate-oval, highest just before middle, dark-brown. Foramen basolateral. Hypandrium light-brown, midventral, elongate, narrow. Aedeagus slim, simple. Two epandrial lobes situated closely to one another, each bearing long seta. Short epandrial seta present. Surstylus light-brown, trilobate; ventral and mid lobes almost fused, with short apical setae; dorsal lobe longer than ventral, narrow, with several short setae. Cercus dark-brown, shorter than surstylus, elongate, dorsally setose, with 3 large and 2 small apical flattened setae, short ventral protuberance and long narrow sclerotised subapical ventral process.
Female unknown.
Length (mm): body without antennae 2.4, antenna 0.9, wing 2.5-2.8/0.9, hypopygium 0.7.
Distribution: South Africa.
Diagnosis. M. africana keys out to M. seksyaevae GRICHANOV 1999a: Fig. 14, differing from the latter species in length ratio of first two segments of fore tarsus (17/48 rather than 15/30) and m-cu/CuA1 wing veins (16/17 rather than 9/18) and morphology of hypopygium. Hypopygium of the new species has some similarity to that in M. capensis CURRAN 1926 and M. hamata PARENT 1936, differing from those species in having long thin epandrial lobes (compare Figs. 9-11).
9. Medetera africana senegalensis GRICHANOV, subspec. nov.
Holotype. G, Senegal, Ziguinchor, 13.VIII.1979, A. PAULY r?c., P[iege] M[alaise] / Coll. Mus. Tervuren [RMCA].
Diagnosis. M. africana senegalensis is almost identical to nominotypical subspecies in any respect including morphology of hypopygium, differing in somewhat smaller size, bluish reflection of face and thorax, mostly black (except for distal 1/3 or 1/4) femora. The habitus is similar to that in M. varitibia PARENT 1935 (male topotypes examined, RINS), which is characterises by midventral projection on surstylus and simple setae on cercus (compare Figs. 9 and 12).
Female unknown.
Length (mm): body without antennae 2.1, wing 2.3.
Distribution: Senegal.
10. Medetera bweza GRICHANOV, spec. nov. (Fig. 13)
Holotype. G, Congo Belge: P.N.A., Tshamugussa (Bweza), 2250 m (Bambous), 10.VIII.1934, G.F. DE WITTE, 526 / P. VANSCHUYTBROECK det. 1951: Saccopheronta hirsuticosta PARENT [RMCA].
Description. Male. Frons and face black, entirely grey pollinose. A row of white postocular setae present. One strong vertical seta laterally on frons. Ocellar tubercle with one pair of strong setae. Ventral postcranium covered with sparse white irregular setae. Face widest under antennae. Antenna with scape and pedicel reddish-yellow; pedicel slightly swollen, with a ring of short apical setulae; first flagellomere brown, small, rounded, as long as high, with distinct apex and short terminal hairs. Arista dorsal (2nd segment broken). Palpus and proboscis short, black, pollinose, with light hairs; palpus with one dark seta.
Thorax bronze-black, grey pollinose, with black setae (mostly broken). Two pairs of strong dorsocentral setae with several hairs anteriorly. Two rows of acrostichals extending to mesonotal flattening. Propleura with 1 strong seta (broken) and 1-2 short white hairs above fore coxa. Scutellum with a pair of strong medial setae and two small lateral hairs.
Legs mostly yellow, with light ciliation, all coxae black-brown with yellow apex, apical segments of tarsi brown. Coxae with white setae and hairs; fore coxa with short hairs and several apical setae; mid and hind coxae each with one external seta. Fore legs without setae. Fore tarsus simple. Length ratio of fore coxa to femur to tibia to tarsus (segments from first to fifth), 33 : 48 : 45 : 19 : 11 : 8 : 6 : 6. Mid tibia with one anterodorsal and one posterodorsal setae at basal 1/4. Mid tarsus broken. Hind legs without long setae. Hind tibia with yellow apical scale only. Hind tarsus with small basoventral tooth. Length ratio of hind coxa to femur to tibia to tarsus (segments from first to fifth), 20 : 55 : 65 : 17 : 27 : 13 : 8 : 7.
Wings hyaline, veins brown. Costa without long hairs. R1 1/3 length of wing. Ratio of part of costa between R2+3 and R4+5 to this between R4+5 and M1+2, 30 : 4. R4+5 and M1+2 distinctly convergent in apical part. Ratio of apical to basal part of M1+2 (from r-m), 90 : 57. Ratio of cross-vein m-cu to maximal distance between R4+5 and M1+2 to apical part of CuA1, 11 : 18 : 28. Lower calypter brownish-yellow, with brownish cilia. Halters light.
Abdomen black, weakly pollinose, with short light setae; 7th segment short. Epandrium black-brown, slightly longer than high. Foramen basolateral. Hypandrium brown, midventral, elongate, narrow, Aedeagus slim, simple. Epandrial lobi reduced to 2 long pedunculate setae positioned closely to one another. Short epandrial seta present. Surstylus yellow, bilobate; ventral lobe with one short and one long apical processes, 2 long flattened and 1 simple apicoventral setae; dorsal lobe long, narrow, with short setae. Cercus brown, shorter than surstylus, elongate, dorsally setose, with 2 apical flattened setae and short subapical ventral process.
Female unknown.
Length (mm): body without antennae 2.0, wing 2.3/0.9, hypopygium 0.7.
Distribution: Congo (Kinshasa).
Diagnosis. M. bweza is closely related to M. edwardsi GRICHANOV 1997a: Fig. 2 and M. norlingi GRICHANOV 1997a: Fig. 3, differing from these species in slightly larger size, somewhat stronger convergent R4+5 and M1+2 veins and hypopygium morphology as follows. Cercus without ventral projection, with 3 curved flattened setae rather than 1-2 simple setae; dorsal lobe of surstylus short rather than long, arising at distal 1/3 or 1/4 and bearing 2 long dorsal setae; ventral process of surstylus with long rather than short apical seta.
11. Medetera calvinia GRICHANOV, spec. nov. (Fig. 14)
Holotype. G, Nieuwoudtville Area, Calvinia district, South West Cape, 14 October 1964, B. & P. STUCKENBERG [NMP].
Paratypes. G, same label; E, Strandfontein coast, West of Van Rhynsdorp, South West Cape, 15-17 October 1964, B. & P. STUCKENBERG [NMP].
Description. Male. Frons and face black, grey-brownish pollinose; clypeus mostly shining. A row of several fine short black postocular setae at the top of eye and a row of white postoculars below present; two lowest postoculars long, brown. One strong vertical seta laterally on frons. Ocellar tubercle with one pair of strong setae and 2 short hairs. Ventral postcranium covered with sparse long white irregular setae. Face widest under antennae, parallel-sided below, 3 times higher than wide in middle. Antenna as long as head height, black; pedicel slightly swollen, with a ring of short apical setulae; first flagellomere small, rounded, with short terminal hairs. Arista subapical. Palpus and proboscis short, black, weakly pollinose, with light hairs; palpus slightly shining, with 1 black seta.
Thorax black, grey pollinose, with black setae. Two pairs of strong dorsocentral setae with row of hair-like setae anteriorly. Two rows of short acrostichals in anterior half of mesonotum. Propleura with 1 strong brownish seta and one short white hair above fore coxa. Scutellum with a pair of strong medial setae and two weak lateral setae, half as long as medial.
Legs black, knees brown. Coxae with light setae; fore coxa with short, shining light hairs; mid and hind coxae each with one fine external seta. Fore legs without strong setae. Fore basitarsus in distal half and 2-4th segments with ventral pile of short erect hairs. Length ratio of fore coxa to femur to tibia to tarsus (segments from first to fifth), 38 : 53 : 50 : 22 : 18 : 10 : 7 : 7. Mid tibia with one anterodorsal, one posterodorsal at basal 1/4 and several very short apical setae. Mid tarsus with apicoventral spinules. Length ratio of mid coxa to femur to tibia to tarsus (segments from first to fifth), 30 : 60 : 67 : 32 : 21 : 16 : 8 : 8. Hind femur without setae. Hind tibia with dorsal swelling at distal 1/4 bearing elongate setulae and one light seta; with one apicoventral seta, several short black spinules covered with light scale posteriorly. Hind basitarsus with elongate apicoventral setula and small posterior apical scale. Length ratio of hind coxa to femur to tibia to tarsus (segments from first to fifth), 18 : 63 : 78 : 18 : 38 : 20 : 10 : 10.
Wings greyish, almost hyaline, veins brown. Costa without long hairs. Ratio of part of costa between R2+3 and R4+5 to this between R4+5 and M1+2, 21 : 6. R4+5 and M1+2 distinctly convergent, R4+5 slightly bowed anteriorly, M1+2 straight. Ratio of apical to basal part of M1+2 (from r-m), 68 : 60. Ratio of cross-vein m-cu to maximal distance between R4+5 and M1+2 to apical part of CuA1, 12 : 17 : 12. Anal vein fold-like. Lower calypter brownish-yellow, with yellow cilia. Halters yellow.
Abdomen flattened dorsoventrally, black, grey pollinose, with short light setae. Epandrium elongate, twice as long as high. Foramen positioned in basal half. Hypandrium arising before middle of epandrium, elongate, narrow. Aedeagus slim, simple. Epandrial lobi reduced to 2 long pedunculate setae positioned closely to one another. Surstylus 1/3 length of epandrium, short and broad, bilobate; ventral lobe with 1 long midventral, 1 thick and short apicoventral, 1 large leaf-like apicodorsal setae; dorsal lobe as long as ventral, with 2 simple apical setae of unequal length and 1 small flattened apical seta. Cercus nearly as long as surstylus, dorsally with short hairs, apically with 3 large flattened setae and subapical ventral projection bearing 3 thick setae.
Female similar to male except lacking male secondary sexual characters. Hind tibia without dorsal setae.
Length (mm): body without antennae 2.1, antenna 0.7, wing 2.3/0.8, hypopygium 0.5.
Distribution: South Africa.
Diagnosis. M. calvinia is close to M. ghesquierei GRICHANOV 1999a: Fig. 11, M. rikhterae GRICHANOV 1997a: Fig. 7, and M. ealensis PARENT 1936, differing by ventral pile on fore tarsus, dorsal swelling on hind tibia, leaf-like setae on surstylus and cercus (compare Figs. 14 and 15 in this paper).
12. Medetera cimbebasia GRICHANOV, spec. nov. (Fig. 16)
Holotype. G, Namibia: L?deritz, Rooiberg, 27°38'S, 16°28'E, 22-24.IX.1997, KIRK-SPRIGGS & MARAIS, Malaise trap sample / Namibian National Insect Collection, National Museum, P.O. Box 1203, Windhoek, Namibia [NMN].
Paratypes. 3GG, 4EE, same labels; G, Namibia: Brandberg, Plateau Valley at: 21°10'46"S, 14°32'52"E, 1950m, 19-21.X.1998, R. BUTLIN & J. ALTRINGHAM, Malaise trap 6; 2GG, Namibia: Brandberg, Pools on Wasserfallfl?che, 21°10'40"S, 14°33'08"E, 2000m, 21-23.X.1998, R. BUTLIN & J. ALTRINGHAM, Malaise trap 8; 1G, 2EE, Namibia: L?deritz, Obib water, 28°00'S, 16°38'E, 19-21.IX.1997, MARAIS & KIRK-SPRIGGS, Malaise trap sample [NMN].
Additional material. 1E, NA99-L03: Namibia: Brandberg, Wasserfallfl?che, 1960m, 21?10'77S, 14?3287E, 07.IV.1999, S. van NOORT & S.G. COMPTON, UV Light trap, overlooking well vegetated Valley below waterfall Bushy Karoo-Namib shrubland; [NMN].
Description. Male. Frons and face dark-green; frons entirely and epistome laterally whitish pollinose; epistome mat-green, clypeus shining blue-green in middle. Postocular setae white, brownish at the top of eye. One strong but short vertical seta laterally on frons. Ocellar tubercle with one pair of strong setae (broken off). Ventral postcranium covered with sparse long white irregular setae. Face 2.3 times higher than wide under antennae. Ratio of height of epistome to its maximal width to its minimal width to height of clypeus to its maximal width, 25 : 15 : 10 : 12 : 15. Antenna short, as long as head height, brown-black; pedicel slightly swollen, with a ring of short apical setulae; first flagellomere small, rounded, with short terminal hairs. Arista apical. Palpus and proboscis short, black, with light hairs; palpus with 1 brown seta.
Thorax bluish-green to blue, whitish pollinose, with black setae. Two pairs of strong dorsocentral setae with anterior seta half as long as posterior; a row of several hairs in front of the 1st dorsocentral seta. Two rows of short acrostichals extending to mesonotal flattening. Propleura with 2 strong white subequal setae above fore coxa. Scutellum with a pair of strong medial setae and two lateral setae, 2/3 as long as medial.
Legs mostly black-brown, coxae black, knees light-brown. Coxae with black setae; fore coxa with short light hairs; mid and hind coxae each with one fine external seta. Fore legs without strong setae. Length ratio of fore coxa to femur to tibia to tarsus (segments from first to fifth), 46 : 62 : 62 : 28 : 35 : 21 : 8 : 9. Mid tibia with one anterodorsal, one posterodorsal at basal 1/4 and several very short apical setae. Length ratio of mid coxa to femur to tibia to tarsus (segments from first to fifth), 25 : 70 : 72 : 45 : 34 : 18 : 7 : 8. Hind femur with row of elongate anteroventral setulae in distal 1/3 and row of elongate dorsal setulae in basal 1/3. Hind tibia slightly swollen at extreme apex, with 1-2 short subapical dorsal setae, short posterodorsal apical hook of 2 curved setae, several black apical spinules covered with brownish scale. Hind basitarsus with small basal tooth posteroventrally. Length ratio of hind coxa to femur to tibia to tarsus (segments from first to fifth), 21 : 70 : 81 : 20 : 48 : 24 : 9 : 9.
Wings hyaline; veins mostly brown, yellow at base. Costa without long hairs. Ratio of part of costa between R2+3 and R4+5 to this between R4+5 and M1+2, 22 : 5. R4+5 and M1+2 distinctly convergent, R4+5 slightly bowed anteriorly. Ratio of apical to basal part of M1+2 (from r-m), 88 : 68. Ratio of cross-vein m-cu to maximal distance between R4+5 and M1+2 to apical part of CuA1, 14 : 19 : 15. Anal vein fold-like. Lower calypter yellow, with yellow cilia. Halters yellow.
Abdomen black, grey pollinose, with short, shining light setae. Epandrium elongate, twice as long as high. Foramen midlateral. Hypandrium arising just before missle of epandrium, elongate, narrow. Aedeagus slim, simple. Epandrial lobe nearly half as wide as surstylus, long, with 2 long flattened setae. Surstylus slightly longer than epandrium; ventral lobe shorter than dorsal, with several short setae at apex; dorsal lobe widened at apex, with 2 apical setae. Cercus shorter than surstylus, dorsally setose, with one small flattened apical seta.
Female similar to male except lacking male secondary sexual characters. Several upper postocular setae dark-brown.
Length (mm): body without antennae 2.5 (male) - 3.0 (female), antenna 0.9, wing 2.7/0.8 (male) - 3.0/1.0 (female), hypopygium 0.8.
Distribution: Namibia.
Etymology. "Cimbebas" is the archaic name for the area between Cape Negro and the Tropic of Capricorn on the southwest African seaboard.
Diagnosis. M. cimbebasia is a sister species to M. vaalensis GRICHANOV, spec. nov., differing from the latter in white setae on coxae, entirely brown-black tibia, elongate-oval epandrium, thick epandrial lobe bearing flattened setae. The new species is also close to M. rikhterae GRICHANOV 1997a: Fig. 7, differing in elongate 2nd segment of fore tarsus, very long cercus and surstylus, flattened setae on epandrial lobe and cercus.
13. Medetera londti GRICHANOV, spec. nov. (Fig. 17)
Holotype. G, South Africa: Natal, Mhlopeni Nature Res., 15 km SE Muden, 2930AB, Coll.: J.G.H. LONDT, Date: 22.XII.1983 [NMP].
Paratype. G, South Africa: Natal, Renegate, SE 3030CD, 11-20.I.1985, J. LONDT, Malaise trap set in riverine bush [NMP].
Additional material. E, S. Africa: Transvaal, 37 km N. Louis Trichardt, Limpopo Valley, 2229DD, Jan. 1975, STUCKENBERG, arid bushveld; 2EE, Capland, Willomore, 20, 29.XII.1925, Dr. BRAUNS [NMP].
Description. Male. Frons and face green-black, brownish pollinose; clypeus mostly shining blue-green. A row of several short brown postocular setae at the top of eye and a row of long white postoculars below present. One strong vertical seta laterally on frons, half as long as ocellar setae. Ventral postcranium covered with sparse long white irregular setae. Face under antennae slightly wider than face at palpi, with weakly concave lateral margins, 2.7 times higher than wide in middle. Antenna as long as head height, black; pedicel slightly swollen, with a ring of short apical setulae; first flagellomere small, rounded, with distinct apex and short terminal hairs. Arista subapical. Palpus and proboscis short, black, weakly shining, with light and dark hairs; palpus with one brown seta.
Thorax bluish-black, grey pollinose, with black setae. Two pairs of strong dorsocentral setae of unequal length; a row of several hairs in front of the 1st dorsocentral seta. Two rows of short acrostichals extending to mesonotal flattening. Propleura with 3 strong white setae of equal length above fore coxa. Scutellum with a pair of strong medial setae and two lateral setae, 2/3 as long as medial.
Legs entirely black-brown. Coxae with white setae; fore coxa with short light hairs and several apical setae; mid and hind coxae each with one fine external seta. Fore legs without strong setae. Fore tarsus simple. Length ratio of fore coxa to femur to tibia to tarsus (segments from first to fifth), 50 : 70 : 72 : 38 : 36 : 20 : 10 : 12. Mid tibia with one anterodorsal, one posterodorsal at basal 1/4 and several very short apical setae. Mid tarsomeres with short apicoventral spinules. Length ratio of mid coxa to femur to tibia to tarsus (segments from first to fifth), 31 : 80 : 88 : 59 : 36 : 20 : 10 : 10. Hind femur with row of several white dorsal setae in basal half, nearly as long as femur diameter. Hind tibia abruptly narrowed at extreme apex, with small brownish apicoventral scale and somewhat larger black apical posterior scale covering extreme base of basitarsus. Hind tarsus simple. Length ratio of hind coxa to femur to tibia to tarsus (segments from first to fifth), 24 : 79 : 100 : 20 : 55 : 28 : 11 : 11.
Wings greyish, almost hyaline; veins brown, yellow at base. Costa without long hairs. Ratio of part of costa between R2+3 and R4+5 to this between R4+5 and M1+2, 23 : 7. R4+5 and M1+2 distinctly convergent, R4+5 slightly bowed anteriorly. Ratio of apical to basal part of M1+2 (from r-m), 109 : 95. Ratio of cross-vein m-cu to maximal distance between R4+5 and M1+2 to apical part of CuA1, 22 : 28 : 15; m-cu weakly convex posteriorly. Anal vein fold-like. Lower calypter yellow, with yellow cilia. Halters yellow.
Abdomen greenish-black, grey pollinose, with short light setae. Epandrium elongate, twice as long as high. Foramen midlateral. Hypandrium midventral, elongate, narrow. Aedeagus slim, simple. Epandrial lobe reduced to short peduncle bearing one long and one short setae. Surstylus short, 2/5 length of epandrium, greatly narrowed in distal half, split at extreme apex, with narrow apical process, several apical and subapical setae as figured. Cercus 2 times shorter than surstylus, elbow-like, dorsally and apically setose, with simple setae.
Female similar to male except lacking male secondary sexual characters.
Length (mm): body without antennae 3.2, antenna 1.0, wing 3.2/1.0, hypopygium 0.94.
Distribution: South Africa.
Etymology. The species is named after one of the collectors, Dr. J.G.H. LONDT.
Diagnosis. M. londti is very close to M. rikhterae GRICHANOV 1997a: Fig. 7, differing in subapical arista, narrow apex of hind tibia, curved and narrowed in distal half surstylus, elbow-like cercus etc.
14. Medetera pallidotiosa GRICHANOV, spec. nov. (Fig. 18)
Holotype. G, Coldstream, 3323Dc, Cape Prov., 25-28 Oct. 1964, STUCKENBERG [NMP].
Paratypes. 2GG, 1E, same label.
Description. Male. Frons and face black, entirely grey pollinose; clypeus shining in middle. A row of black-brown postocular setae present. One strong vertical seta laterally on frons. Ocellar tubercle with one pair of strong setae. Ventral postcranium covered with sparse light irregular setae. Face widest under antennae, narrowest in middle, slightly widened towards palpi. Ratio of height of epistome to its maximal width to height of clypeus to its minimal width, 18 : 12 : 7 : 9. Antenna black, nearly as long as head height; pedicel slightly swollen, with a ring of short apical setulae; first flagellomere small, rounded, as long as high, with short terminal hairs. Arista apical. Palpus and proboscis short, black, pollinose, with light hairs; palpus with one dark seta.
Thorax black, densely grey pollinose, with black setae. Two pairs of strong and 4-5 anterior pairs of very small dorsocentral setae. Two rows of acrostichals not reaching mesonotal flattening. Propleura with 1 strong black seta and 1 short black hair above fore coxa. Scutellum with a pair of strong setae, without lateral hairs.
Legs mostly dirty-yellow, with all coxae black, femora black-brown in basal half, apical segments of tarsi brown. Coxae with black setae and hairs; fore coxa with short hairs and several apical setae; mid and hind coxae each with one external seta. Fore legs without setae. Fore tarsus simple. Length ratio of fore coxa to femur to tibia to tarsus (segments from first to fifth), 30 : 45 : 45 : 20 : 15 : 10 : 8 : 6. Mid tibia with one anterodorsal and one posterodorsal setae at basal 1/4. Mid tarsus simple. Length ratio of mid coxa to femur to tibia to tarsus (segments from first to fifth), 22 : 48 : 55 : 27 : 17 : 11 : 7 : 6. Hind legs without long setae. Hind tibia with posteriorly with several microscopic black spinules covered with apical scale. Hind tarsus simple. Length ratio of hind coxa to femur to tibia to tarsus (segments from first to fifth), 18 : 49 : 68 : 13 : 26 : 14 : 8 : 7.
Wings hyaline, veins brown. Costa without long hairs. R1 1/3 length of wing. Ratio of part of costa between R2+3 and R4+5 to this between R4+5 and M1+2, 20 : 7. R4+5 and M1+2 distinctly convergent in apical part. Ratio of apical to basal part of M1+2 (from r-m), 77 : 50. Ratio of cross-vein m-cu to maximal distance between R4+5 and M1+2 to apical part of CuA1, 12 : 16 : 20 (12 : 16 : 16 in paratypes). Lower calypter brownish-yellow, with brownish, though shining light cilia. Halters yellow.
Abdomen black, weakly pollinose, with short light setae; 7th segment half as long as epandrium. Epandrium twice longer than high, black, with black-brown appendages. Foramen basolateral. Hypandrium brown, basoventral, elongate, narrow. Aedeagus slim, simple. Epandrial lobi reduced to 2 long pedunculate setae positioned closely to one another. Short epandrial seta halfway between pedunculate setae and base of hypandrium present. Surstylus trilobate; ventral lobe broadest, with several long apical setae; middle lobe with subapical process and 1 apical seta; dorsal lobe long, narrow, with 4 long subapical and 1 short apical setae. Cercus nearly as long as surstylus, broad, elongate, dorsally setose, with 2 large apical flattened setae, 2 thick and 3 simple apicoventral setae.
Female similar to male except lacking male secondary sexual characters.
Length (mm): body without antennae 2.0, antenna 0.6, wing 2.1/0.8, hypopygium 0.5.
Distribution: South Africa.
Diagnosis. M. pallidotiosa keys out to M. ghesquierei GRICHANOV, 1999a: Fig. 11, differing in having simple setae on surstylus. The new species is also close to M. nocturna CURRAN 1927 (male determined by O. PARENT was examined, RINS, Fig. 19) and M. praedator CURRAN 1926 (male paratype examined, RMCA, Fig. 20), differing from the 1st species in having simple setae on surstylus, and from the 2nd in absence of long apicodorsal flattened seta on cercus.
Distribution: South Africa.
15. Medetera praedator aequatorialis GRICHANOV, subspec. nov. (Fig. 21)
Holotype. G, [Tanzania:] Makoa, 22-23.II.1952, D.O. Africa Exp. / P. VANSCHUYTBROECK det. 195? Medetera varitibia Par. / R.I.Sc.N.B. I.G. 22942 [RINS].
Paratypes. 2GG, [Congo (Kinshasa):] Kibati (1900 m), 19.I.1934, G.F. DE WITTE, Parc Nat. Albert, 168 / P. VANSCHUYTBROECK det. 1951 Saccopheronta quinta PARENT [RMCA]; 1G, Mus?e du Congo, N. Kivu, X.1933, Kibati (plaine de lave), Dr. de WULF / R. Det. S 2966 / Medetera otiosa PARENT, [det.] O. PARENT [RMCA]; 1G, Urundi: Bururi, alt. 1900 m, 26.II.1948, F. FRAN?OIS / R.I.Sc.N.B. I.G. 24452 [RINS].
Diagnosis. Having two strong scutellar setae and two strong dorsocentral setae, M. praedator aequatorialis is almost identical to nominotypical subspecies in any respect, differing in legs coloration and morphology of hypopygium. M. p. praedator has yellow tibia and broad, rounded at apex middle lobe of surstylus (Fig. 20), whereas M. p. aequatorialis has entirely black legs and narrow pointed middle lobe of surstylus. Nominotypical subspecies is known from South Africa only.
Female unknown.
Length (mm): body without antennae 2.0, wing 2.2/0.9.
Distribution: Tanzania, Burundi, Congo (Kinshasa).
16. Medetera vaalensis GRICHANOV, spec. nov. (Fig. 22)
Holotype. G, S. Africa: Transvaal, nr. Louis Trichardt, Wilies Poort, 2229DD, Jan. 1975, STUCKENBERG, streambank.
Description. Male. Frons greenish-black, brownish pollinose; face metallic, laterally pollinose; epistome mat-green, clypeus shining dark-green. A row of several fine short black postocular setae at the top of eye and a row of long white postoculars below present. One strong vertical seta laterally on frons, as long as distance between ocelli. Ocellar tubercle with one pair of long and strong setae. Ventral postcranium covered with sparse long white irregular setae. Face widest at palpi, narrowed in middle, 3.5 times higher than wide under antennae and 2.3 times higher than maximal width of clypeus. Antenna slightly longer than head height, black-brown; pedicel slightly swollen, with a ring of short apical setulae; first flagellomere small, rounded, with short terminal hairs. Arista apical. Palpus and proboscis short, black, weakly pollinose, with light hairs; palpus mat, with 1 black seta.
Thorax blue-black, grey-white pollinose, with black setae. Four pairs of strong dorsocentral setae gradually decreasing in size anteriorly; each seta 1/2 shorter than preceding one; several hairs in front of the 1st dorsocentral seta. Two rows of short acrostichals extending to mesonotal flattening. Propleura with 3 strong black-brown setae of unequal length above fore coxa. Scutellum with a pair of strong medial setae and two lateral setae, 2/3 as long as medial.
Legs with all coxae and femora black; knees, tibia and basitarsi light-brown; other tarsomeres brown. Coxae with black setae; fore coxa with short, shining light hairs; mid and hind coxae each with one fine external seta. Fore legs without strong setae. Fore tarsus simple. Length ratio of fore coxa to femur to tibia to tarsus (segments from first to fifth), 50 : 68 : 66 : 34 : 42 : 26 : 12 : 9. Mid tibia with one anterodorsal, one posterodorsal at basal 1/4 and several very short apical setae. Length ratio of mid coxa to femur to tibia to tarsus (segments from first to fifth), 35 : 64 : 83 : 55 : 35 : 22 : 10 : 8. Hind femur without long setae, with elongate anterodorsal setulae at base. Hind tibia slightly swollen at extreme apex, with 1-2 short subapical dorsal setae, short posterodorsal apical hook of 2 curved setae, several black apical spinules covered with brownish scale. Hind basitarsus with small basal tooth posteroventrally. Length ratio of hind coxa to femur to tibia to tarsus (segments from first to fifth), 26 : 70 : 95 : 21 : 56 : 28 : 12 : 8.
Wings hyaline; veins mostly brown, yellow at base. Costa without long hairs. Ratio of part of costa between R2+3 and R4+5 to this between R4+5 and M1+2, 24 : 6. R4+5 and M1+2 distinctly convergent, R4+5 slightly bowed anteriorly. Ratio of apical to basal part of M1+2 (from r-m), 94 : 84. Ratio of cross-vein m-cu to maximal distance between R4+5 and M1+2 to apical part of CuA1, 15 : 23 : 18. Anal vein distinct. Lower calypter yellow, with yellow cilia. Halters yellow.
Abdomen blue-black, grey pollinose, with short black, though shining light setae. Hypopygium black, with brown appendages. Epandrium greatly swollen in basal half, slightly longer than high. Foramen midlateral. Hypandrium midventral, elongate, narrow. Aedeagus slim, simple. Epandrial lobe as long and thin process bearing 2 long simple setae at apex. Surstylus shorter than epandrium, narrow; ventral lobe shorter than dorsal, with several short setae at apex; dorsal lobe widened at apex, with 2 apical setae. Cercus shorter than surstylus, dorsally setose, with 3 small flattened apical and 2 simple subapical ventral setae.
Female unknown.
Length (mm): body without antennae 2.7, antenna 1.0, wing 2.8/1.0, hypopygium 1.1.
Distribution: South Africa.
Diagnosis. M. vaalensis is closely related to M. cimbebasia GRICHANOV, spec. nov., differing from the latter in black setae on coxae, light-brown tibia, swollen at base epandrium, narrow epandrial lobe bearing simple setae.
Genus Thrypticus GERSTAECKER, 1864
17. Thrypticus parabellus GRICHANOV, spec. nov. (Fig. 23-24)
Holotype. G, Gillits, Pinetown district, Natal, S. Africa, B. & P. STUCKENBERG [NMP].
Description. Male. Frons and face metallic bluish-green. One strong vertical seta laterally on frons, one fine postvertical seta present. Ocellar tubercle with a pair of strong setae. Face widest under antennae, narrowed downward, clypeus parallel-sided. Ratio of height of epistome to its maximal width to height of clypeus, 18 : 18 : 10. Postocular setae light. Antenna entirely black; pedicel with a ring of short apical setulae; first flagellomere higher than long, transverse-oval, with short hairs. Arista apical, with microscopic hairs. Palpus and proboscis short, black, palpus with one black seta.
Mesonotum flattened in posterior third, metallic bluish-green, with light setae. Pleura metallic green. Four pairs of strong dorsocentral setae decreasing in size anteriorly with short hairs in front of the 1st seta. Acrostichal setae biseriate. Presutural part of mesonotum bearing numerous short irregular hairs. ?One light propleural cilia. Scutellum with a pair of strong setae.
Legs with all coxae and femora metallic green-black; trochanters brown; femora yellow at distal apices; fore and hind tibia mostly brown, yellow at base; mid tibia yellow; fore and hind tarsi entirely black; mid basitarsus mostly brownish; mid tarsi brown-black from tip of basitarsus. Fore coxa with short hairs and several light apical setae; middle coxa with hairs and 2 external setae; hind coxa with one long and 2 somewhat shorter light strong external setae. Fore leg without setae. Mid femur with strong posterior subapical seta. Mid tibia without dorsal setae, with one strong apicoventral seta. Hind femur simple. Hind tibia with short apical setae. 1-4th segments of all tarsi with short apical spinules; 5th segments flattened. Length ratio of fore coxa to femur to tibia to tarsus (segments from first to fifth), 40 : 51 : 53 : 24 : 14 : 8 : 6 : 8. Same ratio for middle leg, 30 : 56 : 58 : 29 : 19 : 10 : 8 : 8. Same ratio for hind leg, 23 : 58 : 74 : 23 : 25 : 15 : 9 : 10.
Wing simple, hyaline, veins mostly brown, yellow at base; posterior wing margin evenly convex; maximum wing-width at the end of CuA1. Costa without long hairs. R1 reaching first third of wing. R2+3 almost straight. Ratio of part of costa between R2+3 and R4+5 to this between R4+5 and M1+2, 25 : 10. Ratio of apical to basal part of M1+2, 98 : 48. R4+5 and M1+2 slightly convex anteriad, almost parallel, inconspicuously convergent in apical part. Ratio of cross-vein m-cu to maximal distance between R4+5 and M1+2 to apical part of CuA1, 15 : 12 : 42. Anal vein fold-like; anal lobe small. Alula undeveloped. Lower calypter yellow, with light cilia. Halter yellow.
Abdomen including epandrium metallic dark-bluish-green, grey pollinose, with light setae. Hypandrium, surstylus and cercus brown-black. Epandrium subtriangular, wide basally, narrowed apically; 7th segment symmetrical, embracing base of epandrium; 8th segment narrow, stick-shaped, lying along left distal margin of 7th segment. One strong curved internal epandrial seta at base of hypandrium. Epandrial lobe elongated, thin, with 2 long setae, one apical and one subapical. Hypandrium arising basoventrally, nearly straight, swollen in basal half (ventral view), long and thin, parallel-sided in distal half, with short constriction in the middle and strong basal melanization beyond constriction. Aedeagus thin, arising from base of epandrium, apically cleft, with broad dorsal and narrow ventral lobes. Surstylus 3/4 as long as epandrium, deflected dorsad, lying conformably with similarly deflected cerci. Surstylus (lateral view) lobate, slightly widened apicad, distally melanised, with strong ventral seta at 2/3; distal third of surstylus convex ventrally, concave dorsally, split at extreme apex, with 1 fine short and 2 microscopic distoventral setae. Cercus oblong, with sparse dorsal and lateral setae and distinct apical digitiform section having rounded apex and bearing 2 rather strong dorsal setae.
Female unknown.
Length (mm): body without antennae 2.4, antenna 0.7, wing 2.4/0.9, hypopygium 0.8.
Distribution: South Africa.
Diagnosis. The new species is very close to T. bellus LOEW, 1869, differing in larger size and following characters. Tibia bicolorate: fore and hind tibia mostly brown, mid tibia entirely yellow; hind coxa with 3 external setae; mid tibia without posterodorsal seta; hypandrium swollen in basal half; surstylus with rather deep incision at extreme apex; cercal apical lobe with rounded apex.
Review of Namibian material
Until recently, very little information was available with respect to the Namibian Dolichopodidae. Twenty species were recorded and described in recent reviews of Afrotropical genera and subfamilies of the family (Grichanov 1996-1999). The majority of species have been collected in the central arid part of the country, being xerophilous and halophilous by nature. Some of these species are widely distributed across semi-deserts or maritime territories of the Old World (Amblypsilopus munroi (Curran, 1924), Hydrophorus praecox Lehmann, 1822, Tachytrechus tessellatus (Macquart, 1842), Thinophilus indigenus Becker, 1902). The remaining are regarded as being endemic to Namibia (Medetera chumakovi Grichanov, 1997, Medetera rikhterae Grichanov, 1997, Thinophilus munroi setiscutellatus Grichanov,1997) or to southern part of Africa (Cemocarus griseatus (Curran, 1926), Hydrophorus vaalensis Parent, 1954, Medetera norlingi Grichanov, 1997, Medetera polleti Grichanov, 1997, Medetera subchevi Grichanov, 1997, Condylostylus imitator Curran, 1924). Namibia as a whole and north-eastern parts of the country (Kwando river basin) has a significant tropical element, with many species common with central Africa or occurring across the continental Afrotropics. To date I have examined about 50 species of the family collected in Namibia, some of which should be described as new taxa, but the real number may reach 100-200 species following a special expedition covering a wider geographical area of the country. First records of a species from Namibia are marked with (!).
SUBFAMILY SCIAPODINAE
Genus Amblypsilopus BIGOT, 1889
1. Amblypsilopus munroi (CURRAN, 1924)
GRICHANOV, 1999b.
=Amblypsilopus parilis PARENT, 1931 (Chrysosoma); GRICHANOV, 1996b:292.
Material examined. 3GG, 14EE, Namibia: Rundu dist., Katara Okawango R., 17°48'56S, 18°53'38E, 20-23.I.1998, KIRK-SPRIGGS & MARAIS, Malaise traps; 1G, 1E, Namibia: Rundu dist., Simanya Okawango River, 17°33'17S, 18°32'30E, 23-24.I.1998, KIRK-SPRIGGS & MARAIS, Malaise traps, riverine forest.
Distribution. South Africa; Namibia, Zimbabwe, Angola, Mozambique, Tanzania, Congo (Kinshasa), Nigeria; Sri Lanka.
Genus Chrysosoma GUERIN-MENEVILLE, 1831
2. Chrysosoma tricrinitum PARENT, 1933
Material examined. 7GG, 2EE, Namibia: West Caprivi Park, Kwando River: Susuwe, 17°45'37S, 23°20'55E, 28.IX.-02.X.1998, A.H. KIRK-SPRIGGS, Malaise trap, dry woodland; 1G, 1E, Namibia: West Caprivi Park, Kwando River: Susuwe, 17°45'37S, 23°20'55E, 26-30.IX.1998, A.H. KIRK-SPRIGGS, Yellow pan, dry woodland; 3GG, 5EE, Namibia: West Caprivi Park, Kwando River: Susuwe, 17°45'37S, 23°20'55E, 26-29.IX.1998, A.H. KIRK-SPRIGGS, Multicolour pan traps, riverine edge.
Distribution. Congo (Kinshasa), Burundi, Sierra Leone, Nigeria, Tanzania, Mozambique, Malawi, South Africa, Namibia (!).
Genus Condylostylus BIGOT, 1859
3. Condylostylus imitator CURRAN, 1924
Material examined. 1G, Nakatwa, Mudamu Game Reserve, 18°11'S, 23°25'E, 08-13.III.1992, E. MARAIS & M. PUSCH.
Distribution. Zimbabwe, Botswana, Namibia (!).
SUBFAMILY DOLICHOPODINAE
Genus Hercostomus LOEW, 1857
4. Hercostomus argyropus par PARENT, 1934
GRICHANOV, 1999c.
Material examined. 1G, Buffalo Base, West Caprivi Park, 18°08'S, 21°41'E, 03-05.IV.1990, E. MARAIS; 8GG, 37EE, Namibia: West Caprivi Park, Kwando River: Susuwe, 17°45'37S, 23°20'55E, 28.IX.-02.X.1998, A.H. KIRK-SPRIGGS, Malaise trap, dry woodland; 1G, Namibia: West Caprivi Park, Kwando River: Susuwe, 17°45'37S, 23°20'55E, 26-29.IX.1998, A.H. KIRK-SPRIGGS, 5 Orange pan experiment; 1E, Namibia: West Caprivi Park, Kwando River: Susuwe, 17°45'37S, 23°20'55E, 26-29.IX.1998, A.H. KIRK-SPRIGGS, Multicolour pan traps, riverine edge; 1G, Botswana: Third Bridge, 19°14'S, 23°21'E, 10.III.1993, E. MARAIS.
Distribution. Kenya; Tanzania, Congo (Kinshasa), Uganda, Burundi, Angola, Namibia (!), Botswana (!).
Genus Lichtwardtia ENDERLEIN, 1912
5. Lichtwardtia angularis (MACQUART, 1842)
Material examined. 1E, Buffalo Base, West Caprivi Park, 18°08'S, 21°41'E, 03-05.IV.1990, E. MARAIS; 1G, Botswana: Third Bridge, 19°14'S, 23°21'E, 10.III.1993, E. MARAIS.
Distribution. Senegal; Gambia, Nigeria, Gabon, Uganda, Kenya, Tanzania, Congo (Kinshasa), Zambia, South Africa, Swaziland, Mozambique, Botswana (!), Namibia (!).
6. Lichtwardtia fractinervis (PARENT, 1929)
GRICHANOV, 1998b.
Material examined. 1G, Namibia: Rundu dist., Simanya Okawango River, 17°33'17S, 18°32'30E, 23-24.I.1998, KIRK-SPRIGGS & MARAIS, Malaise traps, riverine forest; 1E, Namibia: Rundu dist., Katara Okawango R., 17°48'56S, 18°53'38E, 20-23.I.1998, KIRK-SPRIGGS & MARAIS, Malaise traps.
Distribution. Benin; Ghana, Nigeria, Congo (Kinshasa), Uganda, Angola, Namibia, Malawi.
7. Lichtwardtia sukharevae GRICHANOV, 1998b.
Material examined. 1G, Namibia: West Caprivi Park, Kwando River: Susuwe, 17°45'37S, 23°20'55E, 28.IX.-02.X.1998, A.H. KIRK-SPRIGGS, Malaise trap, dry woodland; 1G, Namibia: West Caprivi Park, Kwando River: Susuwe, 17°45'37S, 23°20'55E, 26-29.IX.1998, A.H. KIRK-SPRIGGS, 5 Orange pan experiment; 1G, Namibia: West Caprivi Park, Kwando River: Susuwe, 17°45'37S, 23°20'55E, 26-29.IX.1998, A.H. KIRK-SPRIGGS, 5 Yellow pan experiment.
Distribution. Botswana, Madagascar, Namibia (!).
Genus Pelastoneurus LOEW, 1861
8. Pelastoneurus ambiguus PARENT, 1934
Material examined. 2GG, Namibia: West Caprivi Park, Kwando River: Susuwe, 17°45'37S, 23°20'55E, 28.IX.-02.X.1998, A.H. KIRK-SPRIGGS, Malaise trap, dry woodland; 1G, 2EE, Namibia: Khorixas district, Huab River, Krone 721, 20°37'09S, 13°57'31E, 23-26.X.1998, KIRK-SPRIGGS & MARAIS, Malaise trap.
Distribution. Ghana; Congo (Kinshasa), Namibia (!).
Genus Tachytrechus HALIDAY in WALKER, 1851
9. Tachytrechus tessellatus (MACQUART, 1842)
GRICHANOV, 1998a:120.
Material examined. 1G, Namibia: Naukluft Pk., Tsams Ost spring, 24°14'45S, 16°06'17E, 26-27.IX.1997, KIRK-SPRIGGS & MARAIS, Malaise trap; 1G, Namibia: Khorixas district, Huab River, Krone 721, 20°37'09S, 13°57'31E, 23-26.X.1998, KIRK-SPRIGGS & MARAIS, Malaise trap.
Distribution. Senegal; Gambia, Nigeria, Congo (Kinshasa), Burundi, Tanzania, Aldabra, Ethiopia, South Africa, Swaziland, Malawi, Angola, Namibia, Botswana, Mozambique, Madagascar, Mauritius, Seychelles, Socotra, Egypt; Oriental Region, New Caledonia.
SUBFAMILY MEDETERINAE
Genus Corindia BICKEL, 1986
10. Corindia saegeri GRICHANOV, 1998d
Material examined. 1G, Namibia: West Caprivi Park, Kwando River: Susuwe,
17°45'37S, 23°20'55E, 26-30.IX.1998, A.H. KIRK-SPRIGGS, Riparian
Winkler extract.
Distribution. Congo (Kinshasa), Gabon, Namibia (!).
Genus Medetera FISCHER VON WALDHEIM, 1819
11. Medetera capensis CURRAN, 1926
Material examined. 6GG, 2EE, Namibia: Brandberg, Wasserfallf?che at: 21°13'05S, 14°31'01E, 1980 m, 10-12.XI.1998, A.H. KIRK-SPRIGGS, Malaise traps river bed; 1G, 1E, Namibia: Brandberg, Wasserfallflache at: 21°10'42"S, 14°32'55"E, 2000m, 22.X.1998, R. BUTLIN & J. ALTRINGHAM, at light 1900-0700; 1G, Namibia: Brandberg, Plateau Valley at: 21°10'46"S, 14°32'52"E, 1950m, 21-23.X.1998, R. BUTLIN & J. ALTRINGHAM, Malaise trap 7; 1E, Namibia: Brandberg, Pools on Wasserfallflache, 21°10'40"S, 14°33'08"E, 2000m, 21-23.X.1998, R. BUTLIN & J. ALTRINGHAM, Malaise trap 8; 1E, Namibia: L?deritz dist., 8 km W Rosh Pinah, 27°59'28S, 16°39'14E, 10-26.VIII.1998, KIRK-SPRIGGS & MARAIS, Malaise trap sample; 1G, Namibia: Rundu dist., Halili, 17°52'58S, 19°26'49E, 20.I.1998, KIRK-SPRIGGS & MARAIS, light trap, primary forest.
Distribution: South Africa; Namibia (!).
12. Medetera chumakovi GRICHANOV, 1997a
GRICHANOV, 1997a:183.
Distribution. Namibia.
13. Medetera lvovskii GRICHANOV, 1999
Material examined. 1G, 10 km NE Dikweya, Kavango, 17°41'S, 18°32'E, 14-27.I.1993, E. MARAIS, Pres. pitf. trap; 1E, Chamwaala, Ovambo, 17°25'S, 16°03'E, 21.I.1993, E. MARAIS; 1G, 1E, Etudilondjaba, Ovambo, 17°36'S, 17°36'E, 14-27.I.1993, E. MARAIS, Pres. pitf. trap; 2GG, 1E, Edimba, Ovambo, 17°28'S, 16°23'E, 14-26.I.1993, E. MARAIS, Pres. pitf. trap; 2GG, 1E, Namibia: West Caprivi Park, Kwando River: Susuwe, 17°45'37S, 23°20'55E, 26-30.IX.1998, A.H. KIRK-SPRIGGS, Yellow pan dry woodland.
Distribution. Congo (Kinshasa); Namibia (!).
14. Medetera norlingi GRICHANOV, 1997a
GRICHANOV, 1997a:176.
Material examined. 1E, Waterberg Plateau, Park Restcamp, 20°30'S, 17°14'E, 09-13.IV.1993, Malaise trap; 1E, Namibia: Hereroland East, Eiseb River at: 20°39'S, 20°05'E, 18.XI-15.XII.1988, M. Paxton, E. MARAIS, Pres. pitf. trap; 5EE, Namibia: Rundu dist., 1 km S of Katara, 17°50'25S, 18°54'26E, 22-23.I.1998, KIRK-SPRIGGS & MARAIS, Malaise traps, primary forest; 7EE, Namibia: Rundu dist., Simanya Okawango River, 17°33'17S, 18°32'30E, 23-24.I.1998, KIRK-SPRIGGS & MARAIS, Malaise traps, riverine forest; 1G, 2EE, Namibia: West Caprivi Park, Kwando River: Susuwe, 17°45'37S, 23°20'55E, 28.IX.-02.X.1998, A.H. KIRK-SPRIGGS, Malaise trap, dry woodland; 2EE, Leeupan, Kaudom Game reserve, 18°40'S, 20°52'E, 12-14.I.1991, E. MARAIS.
Distribution. Namibia, Botswana, Angola, South Africa.
15. Medetera normalis CURRAN, 1924
GRICHANOV, 1997a:180.
Material examined. 1E, Namibia: Mariental district, Viljoenskroon, 507, 25°10'S, 19°58'E, 29.III-05.VI.1998, G. Olivier, Malaise traps; 1E, Botswana: Third Bridge, 19°14'S, 23°21'E, 10.III.1993, E. MARAIS; 2EE, Ogongo agric. col., Ovamboland, SE 1715 Cb, IX-26.X.1993, 26.X-01.XII.1993, Pres. pitf. traps; 1E, CDM Camp: Tsumkwe, Bushmanland, SE 1920 Cb, V.1993, pittraps, S.V. Green; 1G, Buffalo Base, West Caprivi Park, 18°08'S, 21°41'E, 03-05.IV.1990, E. MARAIS; 5GG, 15EE, Namibia: West Caprivi Park, Kwando River: Susuwe, 17°45'37S, 23°20'55E, 28.IX.-02.X.1998, A.H. KIRK-SPRIGGS, Malaise trap, dry woodland; 2EE, Namibia: West Caprivi Park, Kwando River: Susuwe, 17°45'37S, 23°20'55E, 26-29.IX.1998, A.H. KIRK-SPRIGGS, Multicolour pan traps, riverine edge; 1E, Windhoek, 22°34'S, 17°05'E, 27.II.1996, coll. J. Fish; 1E, Namibia: L?deritz, Rooiberg, 27°38'S, 16°28'E, 22-24.IX.1997, KIRK-SPRIGGS & MARAIS, Malaise trap sample; 1E, Namibia: Naukluft Pk., Tsams Ost spring, 24°14'45S, 16°06'17E, 26-27.IX.1997, KIRK-SPRIGGS & MARAIS, Malaise trap.
Distribution. South Africa, Namibia, Botswana, Burundi, Tanzania, Congo (Kinshasa), Congo (Brazzaville), Ghana.
16. Medetera polleti GRICHANOV, 1997a
GRICHANOV, 1997a:179.
Distribution. Namibia, Botswana.
17. Medetera pseudotiosa GRICHANOV, 1999
GRICHANOV, 1999a.
Distribution. Namibia, Congo (Kinshasa).
18. Medetera rikhterae GRICHANOV, 1997a
GRICHANOV, 1997a:180.
Material examined. 1G, Namibia: Opuwa dist., Okazawana, 18°25'S, 13°47'E, 31.III.1996, E. MARAIS & A.H. KIRK-SPRIGGS.
Distribution. Namibia.
19. Medetera simplicis CURRAN, 1924
GRICHANOV, 1997a:187.
Distribution. South Africa, Namibia, Congo (Kinshasa).
20. Medetera subchevi GRICHANOV, 1997a
GRICHANOV, 1997a:177.
1G,1E, Namibia: Rundu dist., 1 km S of Katara, 17°50'25S, 18°54'26E, 22-23.I.1998, KIRK-SPRIGGS & MARAIS, Malaise traps, primary forest; 5EE, Namibia: Rundu dist., Simanya Okawango River, 17°33'17S, 18°32'30E, 23-24.I.1998, KIRK-SPRIGGS & MARAIS, Malaise traps, riverine forest; 1E, Namibia: Rundu dist., Katara Okawango R., 17°48'56S, 18°53'38E, 20-23.I.1998, KIRK-SPRIGGS & MARAIS, Malaise traps; 1E, Namibia: Mariental district, Viljoenskroon, 507, 26°08'39S, 19°57'11E, 7-9.II.1998, KIRK-SPRIGGS & MARAIS, Malaise traps; 2GG, 15EE, Namibia: West Caprivi Park, Kwando River: Susuwe, 17°45'37S, 23°20'55E, 28.IX.-02.X.1998, A.H. KIRK-SPRIGGS, Malaise trap, dry woodland; 12GE, Namibia: Mariental district, Viljoenskroon, 507, 25°10S, 19°58E, 08-29.III.1998, 29.III-05.VI.1998, 26.IV-02.V.1998, 19-25.XI.1998, 26.XI-02.XII.1998, G. OLIVIER, Malaise traps; 1E, Ohamwaala, Ovambo, 17°25S, 16°03E, 23.I.1993, E. MARAIS; 10GG, 14EE, Botswana: Third Bridge, 19°14S, 23°21E, 10.III.1993, E. MARAIS; 2GG, 1E, CDM Camp: Tsumkwe, Bushmanland, SE 1920 Cb, V.1993, pittraps, S.V. GREEN; 1E, Odila River at Ovambo, 17°29S, 16°51E, 23.I.1993, E. MARAIS, Malaise traps; 4EE, Kaudom Camp, Kaudom Game reserve, 18°30S, 20°44E, 12-15.II.1992, E. MARAIS & P. PUSCH, Light trap.
Distribution. Namibia, South Africa, Botswana.
Genus Thrypticus GERSTAECKER, 1864
21. Thrypticus kataevi GRICHANOV, 1998e
Material examined. 1G, 1E, NA99-L01: Namibia: Brandberg, Messum Valley 700m 21?1329S, 14?3098E, 03.IV.1999, S. van NOORT & S.G. COMPTON, UV Light trap, sparsely vegetated river Valley Bushy Karoo-Namib shrubland.
Distribution. Congo (Kinshasa), Cameroon, Kenya, Swaziland, Namibia (!).
SUBFAMILY HYDROPHORINAE
Genus Cemocarus MEUFFELS & GROOTAERT, 1984
22. Cemocarus griseatus (CURRAN, 1926)
GRICHANOV, 1997b:155.
Material examined. 4GG, Namibia: L?deritz dist., Van Reenenbaai at: 27°24'19S, 15°12'42E, 3.IX.1998, KIRK-SPRIGGS, Sampling station 63, 64 / National Museum of Namibia, Marine littoral survey 1998; 1E, Namibia: L?deritz dist., Orangemund at: 28°37'16S, 16°26'08E, 25.VIII.1998, KIRK-SPRIGGS, Sampling station 62 / National Museum of Namibia, Marine littoral survey 1998.
Distribution. South Africa, Namibia.
Genus Hydrophorus FALL?N, 1823
23. Hydrophorus hydrophylax PARENT, 1939
DYTE & SMITH 1980:455.
Distribution. Uganda, ?Namibia.
24. Hydrophorus jeanneli PARENT, 1938
DYTE & SMITH 1980:455.
Distribution. Kenya, Zaire, Tanzania, ?Namibia.
25. Hydrophorus praecox (LEHMANN, 1822)
GRICHANOV, 1997b:151.
Material examined. 51GG, 127EE, NA99-L01: Namibia: Brandberg, Messum Valley 700m 21?1329S, 14?3098E, 03.IV.1999, S. van NOORT & S.G. COMPTON, UV Light trap, sparsely vegetated river Valley Bushy Karoo-Namib shrubland; 3GG, 2EE, NA99-L03: Namibia: Brandberg, Wasserfallfl?che, 1960m, 21?1077S, 14?3287E, 07.IV.1999, S. van NOORT & S.G. COMPTON, UV Light trap, overlooking well vegetated Valley below waterfall Bushy Karoo-Namib shrubland; 1G, NA99-L02: Namibia: Brandberg, Wasserfallfl?che, 2000m, 21?1076S, 14?3316E, 06.IV.1999, S. van NOORT & S.G. COMPTON, UV Light trap, open plain with grasses & shrubs Bushy Karoo-Namib shrubland; 1G, NA99-M01: Namibia: Brandberg, Messum Valley 700m 21?1329S, 14?3098E, 02-05.IV.1999 S. van NOORT & S.G. COMPTON, Malaise trap, Bushy Karoo-Namib shrubland; 1E, NA99-M02: Namibia: Brandberg, Messum Valley, 700m, 21?1329S, 14?3098E, 02-05.IV.1999, S. van NOORT & S.G. COMPTON, Malaise trap, Bushy Karoo-Namib shrubland; 1E, NA99-L04: Namibia: Brandberg, K?nigstein, 2470m, 21?0900S, 14? 3451E, 11.IV.1999, S. van NOORT & S.G. COMPTON, UV Light trap, saddle below peak with grasses & shrubs Bushy Karoo-Namib shrubland; 1E, Namibia: Naukluft Pk., Tsams Ost spring, 24°14'45S, 16°06'17E, 26-27.IX.1997, KIRK-SPRIGGS & MARAIS, Malaise trap; 3GG, 6EE, Namibia: L?deritz, Obib water, 28°00'S, 16°38'E, 19-21.IX.1997, MARAIS & KIRK-SPRIGGS, Malaise trap sample; 1G, Bogenfels Area, 27°26S, 15°24'E, Diamond area 1, 11/12 April 1986, J. IRISH; 6EE, Gobabeb, Namib Naukluft Park, 23°24S, 15°03'E, VII.1988, VII.1989, IX.1989, Light trap: Kuiseb; 15GE, Gobabeb, Namib.-Naukluft Park, 23°24S, 15°03'E, XII.1988, M. NEL, Light trap in Kuiseb River; 5GG, 1E, State Museum Windhoek, SMI 2210, Panner HC-7, Rossing Mine Survey, SE 2214 B6, 6/6/84, C. MEYER; 1G, 1E, State Museum Windhoek, SMI 423, Diptera, Okau Sontair Munutum R., 12.5 km inland SE 1812 HC, Skeleton Coast Park, 22 Jan. 83, BETHUNA, DAY; 2GG, 1E, State Museum Windhoek, SMI 422, Diptera, Salt seep, 2 km inland Sechomib R., SE 1812 Cb, Skeleton Coast Park, 22 Jan. 83, BETHUNA, DAY; 1E, State Museum Windhoek, Diptera adult, Hoanib River Khowarib Schlucht, SE 1913 Bd, 11 April 1991, B.A. CURTIS; 1E, State Museum Windhoek, SMI 707, Fish Pond, R?ssing, 3/7/84, C. MEYER; 1E, State Museum Windhoek, SMI 418, Diptera [...] Skeleton Coast Park, 21 Jan. 83, BETHUNA, DAY; 1G, 1E, Boulder Pool 3, R?ssing Survey, SE 2214Db, 04.VI.1984, C. MEYER SMI 2207; 1G, 10 km S Rundu, Kavango, 18°00S, 19°41E, 12.I.1993, E. MARAIS, Light trap; 1G, Tuguva, Kavango, 17°26S, 18°27E, 13.I.1993, E. MARAIS, Light trap; 25GE, Ekuma R. M., Etosha, Pan, Outjo, S.M.N. 22851; 1E, Epembe, Kaokoland, 17°34S, 13°32'E, 5-11.XI.1989, C.S. ROBERTS; 1E, S.M.N. 7535, Okankuejo, Etosha Pan, Outjo, March 1972, H. EBEDES; 1G, 2EE, Omuulu, Ovambo, 17°30S, 16°10E, 14.I.1993, E. MARAIS, at light; 1G, 1E, Orupembe, Kaokoland, 18°10S, 12°34E, 02.V.1991, E. MARAIS.
Distribution. South Africa, Namibia, Angola, Botswana, South Arabia, Ethiopia, Mauritania, Gambia, Nigeria, Kenya, Tanzania, Mauritius, Rodriguez, Aldabra, St. Helena, Cape Verde Islands, Canary Islands; Palearctic, Oriental Regions, Australia, New Zealand.
26. Hydrophorus vaalensis PARENT, 1954
GRICHANOV, 1997b:153.
Distribution. South Africa; ?Namibia, ?Zimbabwe.
Genus Thinophilus WAHLBERG, 1844
27. Thinophilus bipunctatus CURRAN, 1926
Material examined. 2GG, 4EE, Kunene Mouth, Skeleton Coast, 17°16S, 11°47E, 20-22.IV.1994, E. MARAIS, Malaise trap, Yellow tray; 1E, Namibia: Khorixas district, Huab River, Krone 721, 20°37'09S, 13°57'31E, 23-26.X.1998, KIRK-SPRIGGS & MARAIS, Malaise trap.
Distribution. South Africa; Congo (Kinshasa), Namibia (!).
28. Thinophilus indigenus BECKER, 1902
GRICHANOV, 1997c:137.
Distribution. Egypt; Mongolia, Iran, Turkey, Algeria, Cape Verde Is., Ethiopia, South Yemen, Nigeria, Benin, Ghana, Zaire, Tanzania, Madagascar, Angola, Namibia, South Africa, Swaziland; India, Nepal, Malaya, Borneo, Philippines, Taiwan.
29. Thinophilus munroi setiscutellatus GRICHANOV, 1997c
GRICHANOV, 1997c:141.
Material examined. 2EE, Bogenfels Area, 27°26S, 15°24'E, Diamond area 1, 11/12 April 1986, J. IRISH.
Distribution. Namibia.
30. Thinophilus palpatus PARENT, 1930
GRICHANOV, 1997C:144.
Distribution. Ethiopia, Nigeria, Angola, Namibia, Botswana.
31. Thinophilus prudens CURRAN, 1926
Material examined. 1G, Namibia: Khorixas district, Huab River, Krone 721, 20°37'09S, 13°57'31E, 23-26.X.1998, KIRK-SPRIGGS & MARAIS, Malaise trap.
Distribution. South Africa, Namibia (!), Angola, Ghana, Zaire, Senegal.
SUBFAMILY DIAPHORINAE
Genus Asyndetus LOEW, 1869
32. Asyndetus virgatus CURRAN, 1926
Material examined. 2EE, Namibia: Omaruru District Ugab R., 2 km W Brandberg, Wes, 20°58'05"S, 14°06'36"E, 22-24.X.1998, A.H. KIRK-SPRIGGS & E. MARAIS, Malaise trap sample; 1E, NA99-Y88: Namibia: Brandberg, Hungorob Valley 1180m, 21?1140S, 14?3169E, 05-16.IV.1999 S. van NOORT & S.G. COMPTON, Yellow Pan trap, Bushy Karoo-Namib shrubland; 5GG, 5EE, Namibia: West Caprivi Park, Kwando River: Susuwe, 17°45'37S, 23°20'55E, 28.IX.-02.X.1998, A.H. KIRK-SPRIGGS, Malaise trap, dry woodland; 2GG, 1E, Namibia: West Caprivi Park, Kwando River: Susuwe, 17°45'37S, 23°20'55E, 26-29.IX.1998, A.H. KIRK-SPRIGGS, 5 Yellow pan experiment; 6GG, 9EE, Namibia: West Caprivi Park, Kwando River: Susuwe, 17°45'37S, 23°20'55E, 26-30.IX.1998, A.H. KIRK-SPRIGGS, Yellow pan, dry woodland; 1G, 2EE, Namibia: West Caprivi Park, Kwando River: Susuwe, 17°45'37S, 23°20'55E, 26-29.IX.1998, A.H. KIRK-SPRIGGS, Multicolour pan traps, riverine edge; 1G, Namibia: West Caprivi Park, Kwando River: Susuwe, 17°45'37S, 23°20'55E, 26-29.IX.1998, A.H. KIRK-SPRIGGS, 5 White pan experiment; 1G, Leeupan, Kaudom Game reserve, 18°40S, 20°52E, 12-14.I.1991, E. MARAIS.
Distribution. South Africa, Namibia (!).
Genus Chrysotus MEIGEN, 1824
33. Chrysotus inconspicuus LOEW, 1860
Material examined. 1E, NA99-L01: Namibia: Brandberg, Messum Valley, 700m, 21?1329S, 14?3098E, 03.IV.1999, S. van NOORT & S.G. COMPTON, UV Light trap, sparsely vegetated river Valley Bushy Karoo-Namib shrubland; 10EE, Namibia: West Caprivi Park, Kwando River: Susuwe, 17°45'37S, 23°20'55E, 28.IX.-02.X.1998, A.H. KIRK-SPRIGGS, Malaise trap, dry woodland; 1G, 2EE, Namibia: West Caprivi Park, Kwando River: Susuwe, 17°45'37S, 23°20'55E, 26-29.IX.1998, A.H. KIRK-SPRIGGS, 5 Yellow pan experiment; 2EE, Namibia: West Caprivi Park, Kwando River: Susuwe, 17°45'37S, 23°20'55E, 26-29.IX.1998, A.H. KIRK-SPRIGGS, 5 White pan experiment.
Distribution. South Africa; Kenya, Namibia (!).
Genus Cryptophleps LICHTWARDT, 1898
34. Cryptophleps spec. aff. rothii COUTURIER, 1978
Material examined. 1E, Namibia: Omaruru District Ugab R., 2 km W Brandberg, Wes, 20°58'05"S, 14°06'36"E, 22-24.X.1998, A.H. KIRK-SPRIGGS & E. MARAIS, Malaise trap sample.
Distribution. Ivory Coast; Namibia (!).
Genus Diaphorus MEIGEN, 1824
35. Diaphorus lawrencei CURRAN, 1926
Material examined. 1E, Namibia: West Caprivi Park, Kwando River: Susuwe, 17°45'37S, 23°20'55E, 28.IX.-02.X.1998, A.H. KIRK-SPRIGGS, Malaise trap, dry woodland.
Distribution. Mozambique; Chad, Ghana, Congo (Kinshasa), Malawi, South Africa, Namibia (!).
Genus Trigonocera BECKER, 1902
36. Trigonocera africana NAGLIS, 1999
NAGLIS, 1999:333.
Distribution. Zambia, Namibia.
SUBFAMILY SYMPYCNINAE
Genus Campsicnemus HALIDAY in WALKER, 1851
37. Campsicnemus caffer CURRAN, 1926
Material examined. 3GG, 3EE, NA99-L01: Namibia: Brandberg, Messum Valley, 700m, 21?1329S, 14?3098E, 03.IV.1999, S. van NOORT & S.G. COMPTON, UV Light trap, sparsely vegetated river Valley Bushy Karoo-Namib shrubland; 1G, Namibia: West Caprivi Park, Kwando River: Susuwe, 17°45'37S, 23°20'55E, 28.IX.-02.X.1998, A.H. KIRK-SPRIGGS, Malaise trap, dry woodland; 1G, Namibia: West Caprivi Park, Kwando River: Susuwe, 17°45'37S, 23°20'55E, 26-30.IX.1998, A.H. KIRK-SPRIGGS, Yellow pan, dry woodland; 2GG, 1E, Namibia: West Caprivi Park, Kwando River: Susuwe, 17°45'37S, 23°20'55E, 26-29.IX.1998, A.H. KIRK-SPRIGGS, Multicolour pan traps, riverine edge.
Distribution. South Africa; Namibia (!).
Genus Sympycnus LOEW, 1857
38. Sympycnus munroi CURRAN, 1925
Material examined. 1G, NA99-L01: Namibia: Brandberg, Messum Valley, 700m, 21?1329S, 14?3098E, 03.IV.1999, S. van NOORT & S.G. COMPTON, UV Light trap, sparsely vegetated river Valley Bushy Karoo-Namib shrubland; 2GG, Namibia: West Caprivi Park, Kwando River: Susuwe, 17°45'37S, 23°20'55E, 26-30.IX.1998, A.H. KIRK-SPRIGGS, Yellow pan, dry woodland; 6GG, 9EE, Namibia: West Caprivi Park, Kwando River: Susuwe, 17°45'37S, 23°20'55E, 28.IX.-02.X.1998, A.H. KIRK-SPRIGGS, Malaise trap, dry woodland.
Distribution South Africa; Kenya, Congo (Kinshasa), Namibia (!).
KEY TO NAMIBIAN GENERA OF DOLICHOPODIDAE
There are several indeterminable or possibly new species to be described later in the following genera: Acropsilus MIK, 1878, Micromorphus MIK, 1878, Teuchophorus LOEW, 1857.
1. Wing vein M2 present, even if as fold or indication on membrane 2
- Vein M2 absent, without fold or indication on membrane 5
2. Both pairs of scutellar setae long; arista dorsal or dorsoapical; pedicel with long dorsal and ventral setae; 3
-Scutellum usually with one pair of strong setae, lateral setae short, hairlike or absent; other features various 4
3. Males with distoventral hypandrium and modified fore basitarsus, usually with long abdomen, often with disturbed wing venation; wing often with dark brown band; frons of both sexes with raised mound bearing strong vertical seta and several or sometimes numerous hairs Condylostylus
- Males with unmodified fore basitarsus, short abdomen, well developed basoventral hypandrium; M1 gently arched to apex, not recurved basad; frons with vertical seta not arising on setose mound Parentia
4. Arista usually apical on triangular first flagellomere; m-cu sinuous; arista usually long, and more than half body length in females; tibiae often with major setae; male cercus often forked or bearing dorsal tooth Chrysosoma
-Arista usually distinctly dorsal on subrectangular first flagellomere and rarely longer than head width, or if apical or dorsoapical, then always with following characters: tibial chaetotaxy often weak, especially on males; m-cu straight, male cercus simple Amblypsilopus
5. Frons strongly excavated above between eyes; M1+2 with two bends at the middle of distal part; dorsocentral bristles strong in both sexes; arista usually dorsal; strong vertical seta present in both sexes; clypeus adjacent to margin of eyes Mesorhaga
- Frons seldom strongly excavated, M1+2 usually straight or evenly curved, with only one bend, sometimes broken or disrupted at the middle of distal part 6
6. Costa of wing ending at tip of R2+3; M1+2 weak or broken near middle of distal part 7
- Costa of wing extending to tip of M1+2; M1+2 never weaker near middle
of distal part
8
7. Male hypopygium usually with strong macrochaetae; acrostichals usually
present
Asyndetus
- Male hypopygium without strong macrochaetae; acrostichals absent or microscopic Cryptophleps
8. Scape with hairs above 9
- Scape bare above 12
9. Hind basitarsus with distinct bristle above; wing vein M1+2 broken in middle of distal part, joining costal vein just before wing tip, having two stublike veins; R4+5 and distal part of M1+2 nearly parallel Lichtwardtia
- Hind basitarsus without bristles above; R4+5 and distal part of M1+2 usually converging 10
10. Several strong anterodorsal setae in apical half of the hind femur in addition to the true anterior subapical seta; face narrowed under antennae and somewhat widened towards clypeus; wing vein M1+2 usually with gentle curvation before the middle of distal part, then running towards R4+5 and reaching costa far before the tip of wing; arista short and bare; first flagellomere usually short and suboval Tachytrechus
-Hind femur usually with one true anterior subapical seta; face regularly narrowed towards clypeus or parallel-sided; wing vein M1+2 either with curvation beyond the middle of distal part or M1+2 reaching costa near the tip of wing; arista often pubescent; first flagellomere usually subtriangular, asymmetric 11
11. Wing vein M1+2 straight or inconspicuously sinuate in basal 1/3 of distal part, usually joining costal vein just before wing tip; R4+5 and M1+2 weakly convergent in distal part; middle and hind femora always with one strong anterior subapical seta positioned just before apex Hercostomus
- Vein M1+2 usually distinctly bent in distal part with strongly convergent R4+5 and M1+2; if R4+5 and M1+2 straight and parallel to each other, then subapical seta positioned at distal third or just behind the middle of hind femora; middle or hind femur sometimes with 2 or more strong anterior subapical setae Pelastoneurus
12. Posterior slope of mesonotum distinctly flattened between dorsocentral setae 13
- Posterior slope of mesonotum not distinctly flattened, usually with a slight transverse depression just before scutellum 16
13. Hind femur with a true anterior subapical seta; arista dorsal; acrostichal setae absent; hypopygium globular, sessile Micromorphus
- Hind femur without a true anterior subapical seta, sometimes with a few longer setae along anteroventral margin near tip, rarely with a row of anterior setae beyond the middle; arista usually apical or subapical; antenna often radially symmetrical from anterior view 14
14. R4+5 and M1+2 convergent, at most subparallel at apex; thorax densely pollinose ................ Medetera
- R4+5 and M1+2 parallel to apex; thorax shining green ................ 15
15. Female oviscapt blade-like, sclerotised, narrow in dorsal view; male surstylus strongly deflexed dorsad, usually lying conformably with similarly deflexed, oblong-shaped cerci ................ Thrypticus
- Female oviscapt soft, male surstylus and cercus usually not deflexed dorsad ................ Corindia
16. Wing cross-vein m-cu distinctly longer than distal part of CuA1 ................ 17
- Wing cross-vein m-cu not distinctly longer than distal part of CuA1 ................ 18
17. First flagellomere trapezoidal, with subapical arista located in dorsoapical excavation; fore femur without strong setae; 6th and 7th male terga well developed ................ Cemocarus
- First flagellomere with apicoventral incision; fore femur thickened, ventrally with strong bristles and spines; at least 7th male tergum greatly reduced ................ Hydrophorus
18. Hind femur without a true anterior subapical seta, sometimes with a few longer anteroventral setae near tip ................ 19
- Hind femur with a true anterior subapical seta ................ 23
19. Male 1st flagellomere at least twice longer than high at base, bud-like, with abruptly drawn-out apex; arista apical or slightly subapical, sometimes inserted in apical incision .................. 20
- Male 1st flagellomere usually not longer than high, reniform, ovate or subtriangular with rounded, rarely acute apex; or if 1st flagellomere much longer than high, then arista distinctly dorsal .................. 21
20. Male eighth tergite with strong and often long macrochaetae .................. Trigonocera
- Male eighth tergite usually with simple hairs, rarely with short setae .................. Chrysotus (part)
21. Male face usually broad, eyes contiguous or distinctly convergent above antennae; middle tibia often with ventral seta; male eighth tergite with strong macrochaetae; female face usually with parallel sides and antennae positioned in middle of head .................. Diaphorus
- Male frons broad, eyes distinctly convergent or contiguous below antennae; middle tibia rarely with ventral seta; male eighth tergite rarely with strong setae; female face usually narrowed ventrad with antennae positioned at upper third of head .................. 22
22. Male hypopygium pedunculate; 7th and 8th segments are often folded, so as hypopygium seems sessile; hypandrium situated before the middle of ventral side of epandrium; female face usually with distinct setae on clypeus .................. Acropsilus
- Male hypopygium sessile; hypandrium situated beyond the middle of ventral side of epandrium; female face without distinct setae on clypeus .................. Chrysotus
23. Face narrow in middle, extending downward .................. Campsicnemus
- Face narrowed gradually or with more or less parallel sides .................. 24
24. Abdomen broad, rather strongly flattened dorsoventrally; 1st flagellomere rounded .................. Thinophilus
- Abdomen not broad and flattened dorsoventrally, usually cylindrical or tapering; 1st flagellomere subtriangular .................. 25
25. Four or five pairs of strong dorsocentral bristles; male wing costa with long and thick stigma beyond R1 .................. Teuchophorus
- Six pairs of dorsocentral bristles; male wing costa without stigma
beyond R1 .................. Sympycnus
Acknowledgements
I am greatly indebted to Dr. Patrick GROOTAERT and Dr. Marc DE MEYER (Brussels) for their kindness in furnishing an opportunity to study the collections of their Museums. I am also sincerely grateful to Dr. David BARRACLOUGH (Pietermaritzburg) and Ashley H. KIRK-SPRIGGS for loaning the material of the Natal and Namibian Museums. The work was carried out in Brussels, in the laboratory of Patrick GROOTAERT (Institut royal des Sciences naturelles de Belgique), and was financially supported by Belgian Office for Scientific, Technical and Cultural Affairs (1999-2000).
Literature
BICKEL, D.J. (1986): Thrypticus and an allied new genus, Corindia from Australia (Diptera: Dolichopodidae). - Records of the Australian Museum 38(3): 135-151; Sydney.
BICKEL, D.J. (1994): The Australian Sciapodinae (Diptera: Dolichopodidae), with a review of the Oriental and Australasian faunas, and a world conspectus of the subfamily. - Records of the Australian Museum, Suppl. 21: 1-39; Sydney.
DYTE, C.E. & SMITH, K.G. (1980): Family Dolichopodidae. In: CROSSKEY, R.W. (ed.): Catalogue of the Diptera of the Afrotropical Region: 443-463; London: British Museum (Natural History).
GRICHANOV, I.Ya. (1996a): Afrotropical species of the genus Condylostylus BIGOT (Diptera: Dolichopodidae). - International Journal of Dipterological Research 7(3): 217-222; St. Petersburg.
GRICHANOV, I.Ya. (1996b): Four new species of the genus Amblypsilopus BIGOT (Diptera: Dolichopodidae) from Tropical Africa and Papua New Guinea. - International Journal of Dipterological Research 7(4): 285-294; St. Petersburg.
GRICHANOV, I.YA. (1997a): Eight new species of Medetera FISCHER VON WALDHEIM (Diptera: Dolichopodidae) from Tropical Africa. - International Journal of Dipterological Research 8(4): 173-179; St. Petersburg.
GRICHANOV, I.Ya. (1997b): A brief review of the Afrotropical fauna of the subfamily Hydrophorinae (Diptera: Dolichopodidae) with description of Cemocarus stuckenbergi spec.n. - International Journal of Dipterological Research 8(3): 149-157; St. Petersburg.
GRICHANOV, I.Ya. (1997c): Notes on Afrotropical and Palearctic species of the genus Thinophilus WAHLBERG (Diptera: Dolichopodidae) with descriptions of new species. - International Journal of Dipterological Research 8(3): 135-147; St. Petersburg.
GRICHANOV, I.Ya. (1998a):. Afrotropical species of the genus Tachytrechus HALIDAY (Diptera: Dolichopodidae). - International Journal of Dipterological Research 9(2): 115-122; St. Petersburg.
GRICHANOV, I.Ya. (1998b): Afrotropical species of the genus Lichtwardtia ENDERLEIN (Diptera: Dolichopodidae). - International Journal of Dipterological Research 9(3): 221-236; St. Petersburg.
GRICHANOV, I.Ya. (1998c): New data on Sciapodinae (Diptera: Dolichopodidae) with a revised catalogue and keys to Afrotropical species - Bulletin de lInstitut royal des Sciences naturelles de Belgique, Entomologie 68: 79-130; Brussels.
GRICHANOV, I.Ya. (1998d): Three new Afrotropical species of the Australian genus Corindia BICKEL (Diptera: Dolichopodidae). - International Journal of Dipterological Research 9(3): 191-194; St. Petersburg.
GRICHANOV, I.Ya. (1998e): Afrotropical species of the genus Thrypticus GERSTAECKER (Diptera: Dolichopodidae). - International Journal of Dipterological Research, 9(3): 213-220; St. Petersburg.
GRICHANOV, I.Ya. (1999a): A brief review of the Afrotropical fauna of the subfamily Medeterinae (Diptera: Dolichopodidae) with descriptions of a new genus and new species. - Bulletin de lInstitut royal des Sciences naturelles de Belgique, Entomologie 69: 87-112; Brussels.
GRICHANOV, I.Ya. (1999b): New species and new records of Afrotropical Sciapodinae (Diptera: Dolichopodidae). - Bulletin de lInstitut royal des Sciences naturelles de Belgique, Entomologie 69: 113-135; Brussels.
GRICHANOV, I.Ya. (1999c): Afrotropical species of the genus Hercostomus LOEW (Diptera: Dolichopodidae). - International Journal of Dipterological Research 10( 1): 7-44; St. Petersburg.
MEUFFELS, H.J.G. & GROOTAERT, P. (1984): Dolichopodidae (Diptera) from Papua New Guinea I: The genus Cymatopus KERTESZ with a discussion on Abatetia MILLER and Cemocarus gen. nov. - Indo-Malayan Zoology 1: 141-158; Rotterdam.
NAGLIS, S. (1999): A new species of Trigonocera Becker (Diptera, Dolichopodidae) from Tropical Africa. - Studia Dipterologica 6(2): 333-335; Halle.
Authors address
Igor Ya. GRICHANOV
All-Russian Institute of Plant Protection, Podbelskogo 3, St. Petersburg,
Pushkin 189620, Russia: e-mail: rytenkov@nat.spb.su
[Remarks under figures]
Figs 1-4, 6, 8, 10, 12, 13, 15-18, 22, 23. Hypopygium, left lateral
view.
Fig 5. Hypopygium, right lateral view.
Figs 9, 11, 14, 19, 20. Apex of hypopygium, left lateral view.
Fig 7. Segments V-VIII of abdomen and hypopygium, left lateral view.
Fig 21. Surstylus, left lateral view.
Fig 24. Hypandrium, ventral view.
1. Mesorhaga kirkspriggsi GRICHANOV, spec. nov.
2. Condylostylus sinclairi GRICHANOV, spec. nov.
3. Condylostylus ulrichi GRICHANOV, spec. nov.
4-5. Parentia asymmetrica GRICHANOV, spec. nov.
6. Corindia demoulini GRICHANOV, spec. nov.
7. Grootaertia irwini GRICHANOV, spec. nov.
8. Grootaertia brevipennis GRICHANOV, spec. nov.
9. Medetera africana africana GRICHANOV, spec. nov.
10. Medetera capensis CURRAN, 1926
11. Medetera hamata PARENT, 1936
12. Medetera varitibia PARENT, 1935
13. Medetera bweza GRICHANOV, spec. nov.
14. Medetera calvinia GRICHANOV, spec. nov.
15. Medetera ealensis PARENT, 1936
16. Medetera cimbebasia GRICHANOV, spec. nov.
17. Medetera londti GRICHANOV, spec. nov.
18. Medetera pallidotiosa GRICHANOV, spec. nov.
19. Medetera nocturna CURRAN, 1927
20. Medetera praedator praedator CURRAN, 1926
21. Medetera praedator aequatorialis GRICHANOV, subspec. nov.
22. Medetera vaalensis GRICHANOV, spec. nov.
23-24. Thrypticus parabellus GRICHANOV, spec. nov.